Subfamily of cricket-like animals
The Phaneropterinae , the sickle-bearing bush crickets or leaf katydids , are a subfamily of insects within the family Tettigoniidae . They may also be known as "false katydids" (to distinguish them from the "true katydids ") or "round-headed katydids".
The name Phaneropterinae is based upon the Old World genus Phaneroptera P. falcata Barbitistini and Odonturini have become brachypterous .
Description
Adult Phaneropterinae species with eggs Eggs of Microcentrum The legs of individuals in this subfamily vary from genus to genus, but, as in nearly all Orthoptera , the posterior (rear) legs are adapted to leaping, and as such are always much longer than other legs. Phaneropterinae are generally well-camouflaged with green and brown colors being most prevalent, but there are exceptions including certain Aganacris Scaphura Batesian mimics of wasps.
The Phaneropterinae are largely arboreal in habitat. The vast majority of species live in shrubs and trees, feeding on leaves and twigs. Some species might potentially cause significant damage, though usually superficial, when present in large numbers, but this is rare; they usually are solitary, unlike meadow grasshoppers , so much so that they seldom come to human notice.
The ovipositor and male genitalia vary according to the genus. The Phaneropterinae differ from other subfamilies of Tettigoniidae (and other Orthoptera) in their oviposition; their eggs are rarely deposited in the earth, but are either glued in double rows to twigs, or are inserted in the edges of leaves.
Taxonomy
As of August 2026 [ update]
Male Acripeza reticulata Acripezini Brunner von Wattenwyl, 1878 – Australia Acripeza Guérin-Méneville, 1832 Ozphyllum Rentz, Su & Ueshima, 2007 Acrometopini
Acrometopa servillea Auth.: Brunner von Wattenwyl, 1878 – Africa, Europe, the Middle East
Acrometopa Fieber, 1853 Altihoratosphaga Hemp, Voje, Heller, Warchalowska-Sliwa & Hemp, 2010 Conchotopoda Karsch, 1887 Horatosphaga Schaum, 1853 Lamecosoma Ragge, 1960 Peronura Karsch, 1889 Peronurella Hemp, 2018 Prosphaga Ragge, 1960 Tenerasphaga Hemp, 2018 Aegimiini Brunner von Wattenwyl, 1878 – Central and South America Aegimia Stål, 1874 Amblycoryphini
Auth.: Brunner von Wattenwyl, 1878 (synonym Amblycoryphae) – Americas, Africa
Amblycorypha insolita Orophus tessellatus
Subtribe Plangiina Cadena-Castañeda, 2015 Madagascarantia Massa, 2017 Monteiroa Karsch, 1889 Paraplangia Heller, 2018 Plangia Stål, 1873 Pseudoplangia Massa, 2014 Agaurella Uvarov, 1939 Amblycorypha Stål, 1873 Eurycoplangiodes Massa, 2020 Eurycorypha Stål, 1873 Orophus Saussure, 1859 Oxygonatium Ragge, 1980 Paraeurycorypha Massa, 2020 Plangiodes Chopard, 1954
Anaulacomerini
Authority: Brunner von Wattenwyl, 1878 – Central and South America
Abrodiaeta Brunner von Wattenwyl, 1891 Anaulacomera Stål, 1873 Grammadera Brunner von Wattenwyl, 1878 Mendesius Piza, 1960 Montealegrezia Cadena-Castañeda, 2012 Pelecynotum Piza, 1967 Phaneropteroides Piza, 1971 Polichnodes Giglio-Tos, 1898 Rostellula Gorochov, 2018 Separatula Gorochov, 2018 Tenellulus Cadena-Castañeda, 2012 Tomeophera Brunner von Wattenwyl, 1878 Viadana Walker, 1869 (incorporates Ctenophlebia Stål, 1873 )
Aniarellini Brunner von Wattenwyl, 1878 – South America
subtribe Aniarellina Brunner von Wattenwyl, 1878 Aniarella Bolívar, 1906 Buzzettium Cadena-Castañeda, 2026 Engonia Brunner von Wattenwyl, 1878 Euxenica Bruner, 1915 Pseudoburgilis Brunner von Wattenwyl, 1878 Xenicola Uvarov, 1940 subtribe Hyperophorina Cadena-Castañeda, 2026 Burgilis Stål, 1873 Coryphoda Brunner von Wattenwyl, 1878 Hyperophora Brunner von Wattenwyl, 1878 Pseudoburgilis Brunner von Wattenwyl, 1878 Tetanina Cadena-Castañeda, 2026 Tetana Brunner von Wattenwyl, 1878
Anisophyini
Auth.: Cadena-Castañeda, 2026; distribution S. America
subtribe Angarina Cadena-Castañeda, 2026 Angara Brunner von Wattenwyl, 1891 Nanoleptopoda Braun, 2011 Parangara Rehn, 1945 subtribe Anisophyina Cadena-Castañeda, 2026 Anisophya Karabag, 1960 Caimanellus Battiston & Fontana, 2024 Cohnia Buzzetti, Fontana & Carotti, 2010 Hellerium Cadena-Castañeda, 2026 Lorieriella Cadena-Castañeda, 2026
Barbitistini
Auth.: Jacobson, 1905 – Europe to central Asia
Male Barbitistes obtusus Isophya
Ancistrura Uvarov, 1921 - monotypic Ancistrura nigrovittata Andreiniimon Capra, 1937 - monotypic Andreiniimon nuptialis Barbitistes Charpentier, 1825 Dasycercodes Bei-Bienko, 1951 - monotypic Dasycercodes iranicus Euconocercus Bei-Bienko, 1950 Isoimon Bei-Bienko, 1954 - monotypic Isoimon riabovi Isophya Brunner von Wattenwyl, 1878 Kurdia Uvarov, 1916 Leptophyes Fieber, 1853 Metaplastes Ramme, 1939 Orthocercodes Bei-Bienko, 1951 - monotypic Orthocercodes zarudnyi Phonochorion Uvarov, 1916 Poecilimon Fischer, 1853 Polysarcus Fieber, 1853
Catoptropterigini
Auth.: Massa, 2016 – Africa
Catoptropteryx Karsch, 1890 Griffinipteryx Massa, 2016 Centroferini Brunner von Wattenwyl, 1878 – South America Centrofera Brunner von Wattenwyl, 1878 Cosmophyllini Brunner von Wattenwyl, 1878 – South America, Madagascar, Australia Dichopetalini
Auth.: Cadena-Castañeda, 2026 - North & central America
Obolopteryx castanea
Acanthorintes Cohn, Swanson & Fontana, 2014 Arachnitus Hebard, 1932 Dichopetala Brunner von Wattenwyl, 1878 Gymnocerca Cohn, Swanson & Fontana, 2014 Mactruchus Cohn, Swanson & Fontana, 2014 Maxium Cadena-Castañeda, 2026 Obolopteryx Cohn, Swanson & Fontana, 2014 Planipollex Cohn, Swanson & Fontana, 2014 Pterodichopetala Buzzetti, Barrientos-Lozano & Rocha-Sánchez, 2010 Rhabdocerca Cohn, Swanson & Fontana, 2014 Tedcohnia Cadena-Castañeda, 2026
Ducetiini
Auth.: Brunner von Wattenwyl, 1878 (now incorporating Elimaeini Brunner von Wattenwyl, 1891 ) – Africa, India, China, Indochina to Australia
Ducetia
Abaxisotima Gorochov, 2005 Agnapha Brunner von Wattenwyl, 1891 Bulbistridulous Xia & Liu, 1991 Ducetia Stål, 1874 Ectadia Brunner von Wattenwyl, 1878 Elimaea Stål, 1874 Kuwayamaea Matsumura & Shiraki, 1908 Noia Walker, 1870 Orthelimaea Karny, 1926 Paraducetia Gorochov & Kang, 2002 Paragnapha Willemse, 1923 Prohimerta Hebard, 1922 Shirakisotima Furukawa, 1963 Subibulbistridulous Shi, 2002
Dysoniini
Dysonia sp.Markia arizae Auth.: Rehn, 1950 – tropical Americas
Genus group Dysonia (temporary)
Genus group Markia Cadena-Castañeda, 2013
Genus group Paraphidniae Cadena-Castañeda, 2016
Genus group not assigned
Hammatofera Brunner von Wattenwyl, 1878 Quiva Hebard, 1927 Yungasacris Rehn, 1950
Dysmorphini Brunner von Wattenwyl, 1878 – Malesia Dysmorpha Brunner von Wattenwyl, 1878 Ectemnini
Auth.: Cadena-Castañeda, 2015 – tropical Americas
Ectemna Brunner von Wattenwyl, 1878 Euthyrrhachis Brunner von Wattenwyl, 1878 Ephippitythini
Auth.: Brunner von Wattenwyl, 1878; distribution: mostly the Australasian realm , Vietnam
Caedicia simplex
Alectoria superba Alectoria Brunner von Wattenwyl, 1879 Caedicia Stål, 1874 Currimundria Rentz, Su & Ueshima, 2008 Diastella Brunner von Wattenwyl, 1878 Dicorypha Krauss, 1902 Ephippitytha Serville, 1838 Kurandoptera Rentz, Su & Ueshima, 2008 Polichne Stål, 1874 Protina Brunner von Wattenwyl, 1879 Symmachis Brunner von Wattenwyl, 1878 Torbia Walker, 1869
Eurypalpini
Authority: Brunner von Wattenwyl, 1878; distribution: Eastern China, Vietnam, Sumatra, Australia
Ceratopompa Karsch, 1890 Zulpha Walker, 1870 Holochlorini
Auth.: Brunner von Wattenwyl, 1878 – Africa, Asia-Pacific
Holochlora japonica Poecilopsyra octoseriata
Ancylecha Serville, 1838 Arantia Stål, 1874 Arnobia Stål, 1876 Calopsyra Brunner von Wattenwyl, 1891 (includes subgenus Parapsyra Carl, 1914 )Casigneta Brunner von Wattenwyl, 1878 Cesasundana Koçak & Kemal, 2009 Dapanera Karsch, 1889 Elbenia Stål, 1876 : includes subgenus Tamdaopteron Gorochov, 2005 Furnia Stål, 1876 Gonatoxia Karsch, 1889 Holochlora Stål, 1873 Leucopodoptera Rentz & Webber, 2003 Liotrachela Brunner von Wattenwyl, 1878 Molpa Walker, 1870 Phaulula Bolívar, 1906 Phygela Stål, 1876 Poecilopsyra Dohrn, 1892 Pseudopsyra Hebard, 1922 Psyrana Uvarov, 1940 Pulchrapsyra Tan, Liu & Ingrisch, 2024 Rectimarginalis Liu & Kang, 2007 Ruidocollaris Liu, 1993 Sinochlora Tinkham, 1945 Stibaroptera Bolívar, 1906 Stictophaula Hebard, 1922 Sympaestria Brunner von Wattenwyl, 1878 Tapiena Bolívar, 1906
Insarini
Auth.: Rehn, & Hebard, 1914 – tropical & subtropical Americas
Arethaea Arethaea Stål, 1876 Brachyinsara Rehn & Hebard, 1914 Insara Walker, 1869 Montezumina Hebard, 1925 Psilinsara Hebard, 1932 Isopserini Brunner von Wattenwyl, 1878 – Eastern Africa, India, Indochina, Malesia Isopsera Brunner von Wattenwyl, 1878 Pelerinus Bolívar, 1906 Pseudopyrrhizia Brunner von Wattenwyl, 1891 Kevaniellini
Auth.: Massa, 2017 – eastern Africa
Kevaniella Chopard, 1954 Letanini
Auth.: Hebard, 1922 – Asia
Himertula Uvarov, 1940 Letana Walker, 1869 Leptoderini Brunner von Wattenwyl, 1878 – Sri Lanka, Malesia Leptoderes Serville, 1838 Mirolliini
Auth.: Brunner von Wattenwyl, 1878 – Asia
Monticolariini Cadena-Castañeda, 2026 - Central & E. Africa Morgeniini
Auth.: Karsch, 1890 - central and western Africa
Mangomaloba Sjöstedt, 1902 Morgenia Karsch, 1890 Odonturini
Odonturella macphersoniAuth.: Brunner von Wattenwyl, 1878 – Iberian peninsula, North Africa
Odontura Rambur, 1838 Odonturella Bolívar, 1900 Pardalotini
Auth.: Brunner von Wattenwyl, 1878 - Africa
Pardalota Brunner von Wattenwyl, 1878 Poecilogramma Karsch, 1887 Percynini
Auth.: Cadena-Castañeda, 2015 – South America
Percyna Grant, 1964 Sictuna Walker, 1869 Phaneropterini
Auth.: Burmeister, 1838 – most continents, especially Africa
Melidia brunneri
Agennis Brunner von Wattenwyl, 1891 - MadagascarDannfeltia Sjöstedt, 1902 Dioncomena Brunner von Wattenwyl, 1878 Dithela Karsch, 1890 Eucatopta Karsch, 1889 Eulioptera Ragge, 1956 Kefalia Hemp, 2019 Melidia Stål, 1876 - AfricaNephoptera Uvarov, 1929 Paraeulioptera Massa, 2020 Parapyrrhicia Brunner von Wattenwyl, 1891 Phaneroptera Serville, 1831 Phanreticula Massa, 2025 Pigalua Massa, 2017 Pleothrix Ragge, 1980 Polichne Stål, 1874 - Australasia Symmetrokarschia Massa, 2015 Symmetroraggea Massa, 2015 Xenodoxus Carl, 1914
Phyllopterini
Female Resecabimus aratus Auth.: Brunner von Wattenwyl, 1878
Phyllopterina Brunner von Wattenwyl, 1878 – Central and South America Apocerycta Brunner von Wattenwyl, 1878 Arota Brunner von Wattenwyl, 1891 Cephalophylloptera Cadena-Castañeda, 2015 Hyperphrona Brunner von Wattenwyl, 1878 Hyperphorina Cadena-Castañeda, 2015 Itarissa Walker, 1869 Julchiella Cadena-Castañeda, 2015 Meneghelia Piza, 1980 Metaprosagoga Vignon, 1930 Phrixa Stål, 1874 Phylloptera Serville, 1831 Resecabimus Cadena-Castañeda, 2015 Uberabina Cadena-Castañeda, 2015 – South America Polichnodes Giglio-Tos, 1898 Uberaba Bruner, 1915
Plangiopsidini
Auth.: Cadena-Castañeda, 2015 – Africa
Plangiola Bolívar, 1906 Plangiopsis Karsch, 1889 Pycnopalpini
Pycnopalpa bicordata Auth.: Cadena-Castañeda, 2014 – Central and South America
Scambophyllini Brunner von Wattenwyl, 1878 – Indochina, Malesia Scambophyllum Brunner von Wattenwyl, 1878 Scaphura nigra Scaphurini Westwood, 1838 – South America Scaphura Kirby, 1825 Scudderiini
Auth.: Brunner von Wattenwyl, 1878 – Americas
Scudderia
Aganacris Walker, 1871 Caroliniella Cadena-Castañeda, 2015 (note : name conflict with Coleopteran)Ceraia Brunner von Wattenwyl, 1891 Ceraiaella Hebard, 1933 Ctenophorema Piza, 1967 Euceraia Hebard, 1927 Harroweria Hebard, 1927 Homotoicha Brunner von Wattenwyl, 1891 Inscudderia Caudell, 1921 Ligocatinus Rehn, 1901 Parascudderia Brunner von Wattenwyl, 1891 Phanerocercus Piza, 1980 Scudderia Stål, 1873 Theudoria Stål, 1874 Vellea Walker, 1869 Zenirella Piza, 1973
Steirodontini
Auth.: Brunner von Wattenwyl, 1878 – Americas ; 3 subtribes:
Microcentrum retinerve Stilpnochlora couloniana Microcentrina
Auth.: Brunner von Wattenwyl, 1878 (previously tribe Microcentrini) – Americas
Acropsis Uvarov, 1939 Apoballa Brunner von Wattenwyl, 1878 Ischyra Brunner von Wattenwyl, 1878 : includes subgenus Anapolisia , Caauara & Hyalipenna Lamprophyllum Hebard, 1924 Lobophyllus Saussure, 1859 Microcentrum Scudder, 1862 : includes subgenus Boroseiyla Petaloptera Saussure, 1859 Philophyllia Stål, 1873 Phoebolampta Brunner von Wattenwyl, 1878 Raggophyllum Nickle, 1967 Tropicophyllum Koçak & Kemal, 2008 : includes subgenus Capanema Tuaia Mendes, Chamorro-Rengifo & Rafael, 2020 : includes subgenus Capiguara Tukunha Mendes & Rafael, 2021 subtribe Steirodontina - subtropical and tropical Americas Cnemidophyllum Rehn, 1917 Coronophyllum Mendes, Sobral & Silva-Neto, 2023 Emsleyfolium Cadena-Castañeda, Mendes & Alves-Oliveira, 2016 Steirodon Serville, 1831 subtribe Stilpnochlorina Gorochov, 2026 Nicklephyllum Cadena-Castañeda, 2016 Problemovapex Gorochov, 2026 Stilpnochlora Stål, 1873 Syntechna Brunner von Wattenwyl, 1878
Taeniomenini Brunner von Wattenwyl, 1878 – Australia Elephantodeta Brunner von Wattenwyl, 1878 Tinzeda Walker, 1869 Trachyzulphini
Trachyzulpha sp.Auth.: Gorochov, 2014 – China, Indochina, Malesia
Trachyzulpha Dohrn, 1892 Trigonocoryphini
Auth.: Bei-Bienko, 1954 – Middle East, India, Malaysia
Cosmozoma Karsch, 1889 Megotoessa Karsch, 1889 Trigonocorypha Stål, 1873 Turpiliini Brunner von Wattenwyl, 1878 – Central America, Caribbean Turpilia Stål, 1874 Turpiliodes Hebard, 1932 Tylopsidini
Auth.: Brunner von Wattenwyl, 1878 – Africa, Europe, Middle East, western Asia
Tylopsis lilifolia Tylopsis Fieber, 1853 Vossiini
Auth.: Cadena-Castañeda, 2015 – Africa, Asia
subtribe Vossiina Cadena-Castañeda, 2015 – Africa Azamia Bolívar, 1906 Vossia Brunner von Wattenwyl, 1891 subtribe Xantiina Cadena-Castañeda, 2015 – Asia Paraxantia Liu & Kang, 2009 – ChinaXantia Brunner von Wattenwyl, 1878 – Borneo
Female Eulophophyllum thaumasium Paracaedicia serrataThese genera have not been placed in a tribe:
Alloducetia Xia & Liu, 1992 Anchispora Brunner von Wattenwyl, 1891 Angustithorax Massa, 2015 Anisotochra Karsch, 1889 Anormalous Liu, 2011 Aphroptera Bolívar, 1902 Astathomima Karny, 1929 Austrodontura Fontana & Buzzetti, 2004 Balneum Piza, 1967 Baryprostha Karsch, 1891 Bertius Piza, 1974 Bongeia Sjöstedt, 1902 Brinckiella Chopard, 1955 Brycoptera Ragge, 1981 Bueacola Sjöstedt, 1912 Chinensis Özdikmen, 2009 Chloroscirtus Saussure & Pictet, 1897 - Americas Choirorhynchus Piza, 1974 Conversifastigia Liu & Kang, 2008 Corycomima Karsch, 1896 Corymeta Brunner von Wattenwyl, 1878 Diastellidea Bolívar, 1902 Dicranopsyra Dohrn, 1892 Ectomoptera Ragge, 1980 Elbeniopsis Dohrn, 1906 Enthephippion Bruner, 1915 Ephippitythoidea Tepper, 1892 Eulophophyllum Hebard, 1922 Euryastes Ragge, 1980 Euxenica Bruner, 1915 Execholyrus Henry, 1940 Gabonella Uvarov, 1940 Gatunella Uvarov, 1940 Gelatopoiidion Zacher, 1909 Gregoryella Uvarov, 1925 Harposcepa Karsch, 1896 Indogneta Ingrisch & Shishodia, 2000 Itokiia Sjöstedt, 1902 Ivensia Bolívar, 1890 Japygophana Carl, 1921 Khaoyaiana Ingrisch, 1990 Ladnea Walker, 1869 Lunidia Hemp, 2010 Macedna Karsch, 1891 Materuana Hemp, 2017 Meruterrana Sjöstedt, 1912 Milititsa Burr, 1900 Miltinobates Sjöstedt, 1902 Mimoscudderia Carl, 1914 Myllocentrum Ragge, 1962 Natricia Walker, 1869 Nesoscirtella Carl, 1914 Niphella Bolívar, 1900 Oxyecous Chopard, 1935 Paracaedicia Brunner von Wattenwyl, 1891 Parapelerinus Liu & Kang, 2008 Paraperopyrrhicia Ebner, 1915 Paraphylloptera Carl, 1914 Parapolichne Bolívar, 1902 Phaneroptila Uvarov, 1957 Physocorypha Karsch, 1896 Platycaedicia Hebard, 1922 Platylyra Scudder, 1898 Polygamus Carl, 1914 Procaedicia Bolívar, 1902 Pronomapyga Rehn, 1914 Pseudomacedna Liu, 2014 Pseudopreussia Hemp, 2017 Puerula Bolívar, 1906 Qinlingea Liu & Kang, 2007 Raggeiella Cadena-Castañeda, 2015 Raggophyllum Nickle, 1967 Sanabria Walker, 1869 Scolocerca Ragge, 1980 Semicarinata Liu & Kang, 2007 Sikoriella Carl, 1914 Stylomolpa Karny, 1926 Sympaestroides Willemse, 1942 Tropidonotacris Chopard, 1954
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