| Two-spotted lady beetle | |
|---|---|
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Clade: | Pancrustacea |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | Polyphaga |
| Infraorder: | Cucujiformia |
| Family: | Coccinellidae |
| Genus: | Adalia |
| Species: | A. bipunctata |
| Binomial name | |
| Adalia bipunctata | |
| Synonyms | |
| |
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Adalia bipunctata, the two-spot ladybird, two-spotted ladybug or two-spotted lady beetle, is a carnivorous beetle of the family Coccinellidae that is found throughout the holarctic region. It is very common in western and central Europe. It is also native to North America, but its population has declined sharply in many states and provinces. It is commonly introduced and imported as a biological control agent.
The two-spotted ladybird was one of the many species originally described by Carl Linnaeus in his 1758 10th edition of Systema Naturae; its original name was Coccinella bipunctata. Its specific name comes from the Latin bi- ("two") and punctata ("spotted").
Adalia bipunctata is a small coccinellid that can feature any one of a large selection of red and black forms. Some forms are similar to Mulsantina picta, but the two white spots on the head of Adalia, in contrast with a large white region or more than two spots, readily distinguish it. Additionally, Adalia is entirely black on the ventral surface with black legs, which helps rule out any other options.[citation needed]
The two-spotted ladybird is highly variable in many parts of its native range. The most familiar form, bipunctata (forma typica), has two black spots on a red base and is common throughout its native range. A melanistic form that is black with four or six red spots is uncommon, but not rare, while the truly melanistic form is exceedingly rare. In North America, the species shows the most variation, with several forms that do not occur elsewhere, including a spotless form, a four-banded form, a form with nine to twelve spots, and a "cross-hatched" form. In addition, there are intermediate forms such as form annulata, but they occur rarely.[failed verification]
Two-spotted lady beetles feed on aphids and other small insects. However, the sterile soldiers within colonies of aphids such as the gall-forming Pemphigus spyrothecae can attempt to protect the aphid colony by fighting this species.[citation needed]

The two-spotted lady beetle's life cycle starts with eggs that are usually laid in clutches. The larva hatches from the egg by biting a hole in it. The larva looks very different from an adult; it has an elongated, grey, soft body with six legs but no wings. The larvae are cannibalistic. A larva goes through four larval stages. As it feeds and grows, it periodically sheds its old skin, allowing further growth. In its last larval stage, the larva is approximately the size of an adult beetle. Once it has eaten enough, the larva attaches itself to a substrate and moults into a pupa. Inside the pupa, the adult develops. Finally, the adult ecloses from the pupa.[citation needed]
In some populations, the majority of the beetles are female. In these populations, 80–90% of the offspring are female. The cause of this anomaly is the presence of symbiotic bacteria living within the gametic cells of the female lady beetles. The bacterium is too large to live in the male gametes (sperm), so the bacterium can be transmitted to the next generation only through female gametes. When it ends up in a male, it will die when the male dies. Therefore, it kills most of the male embryos in the newly laid eggs. These dead embryos then serve as food for their sisters when they emerge from their eggs. This trait is associated with a variety of bacteria (Wolbachia, Rickettsia, and Spiroplasma), which are present in 0–20% of females, depending on the locality.[citation needed]
The two-spot ladybird also carries a sexually transmitted infection in Central and Eastern Europe. This infection is caused by the ectoparasitic mite Coccipolipus hippodamiae, which is transmitted between males and females during copulation. The infection sterilises female two-spot ladybirds, and at certain times of the year, up to 90% of adult two-spots become infected.
A. bipunctata is used as a localised biological control agent against aphids in, for example, greenhouses. The two-spotted lady beetle was introduced into Australia specifically as a biological control agent.