↑(en) Niklas Wahlberg, Jadranka Rota, Michael F. Braby, Naomi E. Pierce et Christopher W. Wheat, «Revised systematics and higher classification of pierid butterflies (Lepidoptera: Pieridae) based on molecular data», Zoologica Scripta, Wiley-Blackwell et Wiley, vol.43, no6, , p.641-650 (ISSN0300-3256 et 1463-6409, DOI10.1111/ZSC.12075)..