↑Christoph D. Schubart, «Reconstruction of phylogenetic relationships within Grapsidae (Crustacea: Brachyura) and comparison of trans-isthmian versus amphi-atlantic gene flow based on mtDNA», Zoologischer Anzeiger, vol.250, , p.472–478 (lire en ligne)
↑Peter K. L. Ng, Danièle Guinot & Peter J. F. Davie, «Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world», Raffles Bulletin of Zoology, vol.17, , p.1–286 (lire en ligne)