Endomychidae, also commonly known as handsome fungus beetles, is a family of coccinelloidbeetles with representatives found in all biogeographic realms. As the common name suggests, members of this family feed on fungi.
Description
They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment.
Taxonomy
The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea.
Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.
This family contains around 120 genera and 1300 species. A list of subfamilies can be seen below:
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