Georgaello

Georgaello
Temporal range: Tithonian
Scientific classification
Kingdom:
Phylum:
Class:
Infraclass:
Genus:
†Georgaello

Greenfield, 2026
Type species
†Georgaello elongata
(Münster, 1836)
Other species
  • †Georgaello wagneri
    (Agassiz, 1843)
Synonyms
Genus synonymy
Species synonymy
  • G. elongata
      • Aellopos elongata
        Münster, 1836
      • Aellopus elongata
        (Münster, 1836)
    G. wagneri
      • Aellopos wagneri
        Agassiz, 1843
      • Aellopus wagneri
        (Agassiz, 1843)
      • Notidanus wagneri
        (Agassiz, 1843)
      • Hexanchus wagneri
        (Agassiz, 1843)

Georgaello is an extinct genus of neoselachians that lived during the Late Jurassic. It contains two species, G. elongata and G. wagneri, which have been found in Germany.

Taxonomy

Aellopos elongata was named by Georg zu Münster in 1836 for a partial holomorphic specimen, a posterior body with dorsal and caudal (and possibly other) fins, from the Kelheim locality in Germany. Aellopos wagneri was named by Louis Agassiz in 1843 for another partial holomorphic specimen, essentially the same parts as A. elongata, apparently from the Solnhofen locality in Germany. The holotypes of both species are housed at the Bayerische Staatssammlung für Paläontologie und Geologie; either of them may or may not be the specimen catalogued as SNSB-BSPG AS VII 1170. Additionally, some vertebrae were later extracted from the A. elongata holotype and possibly taken to the Uniwersytet Wrocławski.

The original genus name is preoccupied by the lepidopteran Aellopos named by Jacob Hübner in 1819. In 1846, Agassiz proposed the unjustified, but available, emendation Aellopus. However, this name is preoccupied by the hemipteran Aellopus named by Johann P. Wolff [de] in 1811. The unavailable, incorrect subsequent spellings Aelopos and Allopos were used by Christoph Giebel in 1846 and Henry W. Fowler in 1911, respectively. A. wagneri was also formerly considered a species of Notidanus or Hexanchus but is not actually assignable to them. Since none of the aforementioned genera can be applied, Georgaello was named as a replacement in 2026.

G. wagneri was the first chondrichthyan recognized from the Solnhofen Limestone, while G. elongata was one of the first named as a distinct genus and species. The former was briefly described as Squalus by Rudolf Wagner in 1829 and the latter was named the same year as Asterodermus platypterus. Both species were redescribed by Johann A. Wagner in 1861, and G. elongata by Carl Hasse [de] in 1882 and Otto M. Reis [de] in 1893.

Classification

The species of Georgaello were previously considered similar to and/or synonymous with several chondrichthyans from the Late Jurassic of Europe, including Pseudorhina acanthoderma, Notidanoides muensteri, Spathobatis bugesiaca, Archaeogracilidens macer, and Aellopobatis bavarica. They have also been compared with extant genera such as Galeorhinus, and Pristis and Pristiophorus.

Currently, the classification of Georgaello cannot be narrowed down past Neoselachii due to a lack of illustrations of the entire holotypes, uncertainty over which museum specimens the holotypes correspond to, and conflicting descriptions of the anatomy. The figured vertebrae of G. elongata have a tectospondylic calcification pattern like those of Pseudorhina spp. and Notidanoides muensteri. The figured dermal denticles match morphotypes found in Asterodermus platypterus, Pseudorhina alifera, Phorcynis catulina, Palaeoscyllium formosum, Paracestracion falcifer, and Macrourogaleus hassei.

Notes

  1. ↑ Other sources have claimed that pelvic and/or pectoral and anal fins are also present in this specimen.
  2. ↑ Other sources have claimed that this specimen originated from the Pappenheim or Eichstätt localities.

References

  1. 1 2 3 4 5 6 7 Greenfield, T. (2026). "Georgaello nom. nov., a replacement name for Aellopos Münster, 1836 (Chondrichthyes, Neoselachii)". Zootaxa. 5886 (4): 589–596. doi:10.11646/zootaxa.5886.4.7.
  2. 1 2 Münster, G.L.F.W.z. (1836). "Capitosaurus in Keupersandstein; Pterodactylus longipes von Solenhofen; Gyrodus multidens in Jurakalk; Aellopus [sic] elongata daselbst; achte Belonostomus-Art; Fische und Caulerpit zu Seefeld; Lebias und Cyprinus in Braunkohle im Fichtelgebirge; vollständiger Placodus gigas; Lumbricarien; Verwechslung sekundärer und tertiärer Schichten zu Solenhofen; Loligo subsagittata; Onychoteuthis speciosa; neues Seestern-Geschlecht von Solenhofen (sonst Phalangites); Squilla antiqua vom Bolca; Insekten von Solenhofen". Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde. 1836: 580–583.
  3. 1 2 3 Agassiz, J.L.R. (1837–1843). Recherches sur les poissons fossiles. Tome III. Neuchâtel, CH & Soleure, CH: Petitpierre & Jent et Gassmann. pp. 376-377. doi:10.5962/bhl.title.4275.
  4. 1 2 Hübner, J. (1816–1826). Verzeichniß bekannter Schmettlinge. Augsburg, DE: Jacob Hübner. p. 131. doi:10.5962/bhl.title.48607.
  5. 1 2 Agassiz, J.L.R. (1846). Nomenclatoris zoologici index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Soleure, CH: Jent et Gassmann. p. 9. doi:10.5962/bhl.title.49761.
  6. 1 2 Wolff, J.P. (1811). "Praefatio editoris". In Wolff, J.F. (ed.). Icones cimicum descriptionibus illustratae. Fasciculus quintus et ultimus. Erlangen, DE: Ioannem Iacobum Palm. pp. iii–v. doi:10.5962/bhl.title.15807.
  7. 1 2 Giebel, C.G.A. (1846). Paläozoologie. Entwurf einer systematischen Darstellung der Fauna der Vorwelt. Merseburg, DE: Nulandt (L. Garcke). p. 188. doi:10.5962/bhl.title.39515.
  8. 1 2 Fowler, H.W. (1911). "A description of the fossil fish remains of the Cretaceous, Eocene and Miocene formations of New Jersey". Geological Survey of New Jersey Bulletin. 4: 1–192. doi:10.5962/bhl.title.63936.
  9. 1 2 3 4 5 Wagner, J.A. (1861). "Monographie der fossilen Fische aus den lithographischen Schiefern Bayerns. Erste Abtheilung: Plakoiden und Pyknodonten". Abhandlungen der Mathemat.-physikalischen Classe der Königlich Bayerischen Akademie der Wissenschaften. 9 (2): 277–352.
  10. 1 2 Fowler, H.W. (1967). "A catalog of world fishes (VI)". Quarterly Journal of the Taiwan Museum. 20 (1–2): 79–148.
  11. 1 2 3 Reis, O.M. (1893). "Untersuchungen über die Petrificirung der Muskulatur". Archiv für Mikroskopische Anatomie. 41: 492–584. doi:10.1007/bf02954332.
  12. 1 2 Wagner, R. (1829). "Nachrichten von der Petrefactensammlung der Königlichen Academie der Wissenschaften in München". Archiv für die gesammte Naturlehre. 16 (1): 71–96.
  13. 1 2 3 4 5 6 Hasse, C. (1882). Das natürliche System der Elasmobranchier auf Grundlage des Baues und der Entwicklung ihrer Wirbelsäule. Eine morphologische und paläontologische Studie. Besonderer Theil. Jena, DE: Gustav Fischer. pp. 104-108 13-14. doi:10.5962/bhl.title.8431.
  14. 1 2 Türtscher, J.; Jambura, P.L.; Villalobos-Segura, E.; López-Romero, F.A.; Underwood, C.J.; Thies, D.; Lauer, B.; Lauer, R.; Kriwet, J. (2024). "Rostral and body shape analyses reveal cryptic diversity of Late Jurassic batomorphs (Chondrichthyes, Elasmobranchii) from Europe". Papers in Palaeontology. 10 (2) e1552. Bibcode:2024PPal...10E1552T. doi:10.1002/spp2.1552. PMC 7615989. PMID 38799546.
  15. ↑ Agassiz, J.L.R. (1835–1843). Recherches sur les poissons fossiles. Tome III. Atlas. Neuchâtel, CH: H. Nicolet. p. 44. doi:10.5962/bhl.title.4275.
  16. ↑ Quenstedt, F.A. (1865–1866). Handbuch der Petrefaktenkunde. Zweite umgearbeitete und vermehrte Auflage. Tübingen, DE: H. Laupp. pp. 205-206.
  17. ↑ Woodward, A.S. (1889). Catalogue of the Fossil Fishes in the British Museum (Natural History), Cromwell Road, S.W. Part I. Containing the Elasmobranchii. London, GB: Trustees of the British Museum. p. 78. doi:10.5962/bhl.title.161983.
  18. ↑ Carvalho, M.R.d.; Kriwet, J.; Thies, D. (2008). "A systematic and anatomical revision of Late Jurassic angelsharks (Chondrichthyes: Squatinidae)". In Arratia, G.; Schultze, H.-P.; Wilson, M.V.H. (eds.). Mesozoic Fishes 4 – Homology and Phylogeny. Munich, DE: Verlag Dr. Friedrich Pfeil. pp. 469–502.
  19. ↑ Kriwet, J.; Klug, S. (2014). "Dental patterns of the stem-group hexanchoid shark, Notidanoides muensteri (Elasmobranchii, Hexanchiformes)". Journal of Vertebrate Paleontology. 34 (6): 1292–1306. Bibcode:2014JVPal..34.1292K. doi:10.1080/02724634.2014.874353.
  20. ↑ Thies, D. (1995). "Placoid scales (Chondrichthyes: Elasmobranchii) from the Late Jurassic (Kimmeridgian) of northern Germany". Journal of Vertebrate Paleontology. 15 (3): 463–481. Bibcode:1995JVPal..15..463T. doi:10.1080/02724634.1995.10011242.