Tobochares
Taxonomy
The genus Tobochares belongs in the subfamily Acidocerinae . It was first described by Short and García in 2007. Since then, as more species are discovered in the seepages of the Guiana Shield Region , there have been two additional papers describing several additional species.
Description
Small beetles (1.5–2.6 mm), yellowish brown, orange brown to dark brown in coloration; eyes entire to emarginate; maxillary palps from short and slender to very short and stout. The elytral punctation and sculpture is highly variable in this genus. There is also variation in the shape of the male genitalia and wing development. A complete diagnosis was presented by Girón and Short.
Habitat
Most Tobochares are considered seepage specialists. One species (Tobochares fusus ) has been also collected in rotten fruits away from water sources.
Species
Tobochares akoerio Girón and Short, 2021 : Suriname
Tobochares arawak Girón and Short, 2021 : Guyana
Tobochares anthonyae Girón and Short, 2021 : Venezuela
Tobochares atures Girón and Short, 2021 : Venezuela
Tobochares benettii Girón and Short, 2021 : Brazil (Amazonas )
Tobochares canaima Girón and Short, 2021 : Venezuela
Tobochares canaliculatus Kohlenberg and Short, 2017 : Venezuela
Tobochares canthus Kohlenberg and Short, 2017 : Venezuela
Tobochares communis Girón and Short, 2021 : Brazil (Amapá , Roraima ), Guyana , Suriname , Venezuela
Tobochares emarginatus Kohlenberg and Short, 2017 : Suriname
Tobochares fusus Girón and Short, 2021 : Brazil (Amapá ), French Guiana
Tobochares goias Girón and Short, 2021 : Brazil (Goiás )
Tobochares kappel Girón and Short, 2021 : Suriname
Tobochares kasikasima Short, 2013 : Suriname
Tobochares kolokoe Girón and Short, 2021 : Suriname
Tobochares kusad Kohlenberg and Short, 2017 : Brazil (Roraima ), Guyana
Tobochares luteomargo Girón and Short, 2021 : Venezuela
Tobochares microps Girón and Short, 2021 : Suriname
Tobochares pallidus Kohlenberg and Short, 2017 : Venezuela
Tobochares pemon Girón and Short, 2021 : Venezuela
Tobochares romanoae Kohlenberg and Short, 2017 : Brazil (Roraima )
Tobochares sipaliwini Short and Kadosoe, 2011 : Brazil (Roraima ), Guyana , Suriname
Tobochares striatus Short, 2013 : Suriname
Tobochares sulcatus Short and García, 2007 : Venezuela
References
1 2 3 Girón, Jennifer C.; Short, Andrew Edward Z. (2021-06-18). "The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species" . ZooKeys (1045): 1– 236. doi :10.3897/zookeys.1045.63810 . ISSN 1313-2970 . PMC 8233300 . PMID 34228772 .
1 2 Short, Andrew Edward Z.; García, Mauricio M. (2007). "Tobochares sulcatus , a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae: Hydrophilini)" . Aquatic Insects . 29 (1): 1– 7. doi :10.1080/01650420701247869 . ISSN 0165-0424 . S2CID 86732116 .
1 2 3 4 5 6 7 8 9 Kohlenberg, Alex T.; Short, Andrew Edward Z. (2017-04-21). "Revision of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)" . ZooKeys (669): 113– 146. doi :10.3897/zookeys.669.11773 . ISSN 1313-2970 . PMC 5523376 . PMID 28769656 .
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Girón, Jennifer C.; Short, Andrew Edward Z. (2021-02-22). "Review of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records" . ZooKeys (1019): 93– 140. doi :10.3897/zookeys.1019.59881 . ISSN 1313-2970 . PMC 7921074 . PMID 33688849 .
1 2 Short, A. E. Z. (2013). "Aquatic beetles of the Grensgebergte and Kasikasima regions, Suriname (Insecta: Coleoptera)". A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname . p. 79. doi :10.1896/054.067.0112 . ISBN 9781934151570 .