(en) John H. Boyle, Zofia A. Kaliszewska, Marianne Espeland, Tamara R. Suderman, Jake Fleming, Alan Heath et Naomi E. Pierce, «Phylogeny of the Aphnaeinae: myrmecophilous African butterflies with carnivorous and herbivorous life histories», Systematic Entomology, Wiley-Blackwell et Wiley, vol.40, no1, , p.169-182 (ISSN0307-6970 et 1365-3113, DOI10.1111/SYEN.12098).