Craspedida is an order of choanoflagellate, with members with an exclusively organic covering. Crown group craspedids (and perhaps crown group choanoflagellates if Acanthoecida arose within Craspedida) appeared 422.78 million years ago. Although a previous study from 2017 recovered the divergence of the crown group choanoflagellates (craspedids) at 786.62 million years.
Extended phylogeny of the Craspedida (Choanomonada). Alexandra Jeuck, Hartmut Arndt and Frank Nitsche, European Journal of Protistology, Volume 50, Issue 4, August 2014, Pages 430–443, doi:10.1016/j.ejop.2014.06.001