It contains 29 described species and 39 subspecies classified in the following five genera:
There are differences in the shape of the forewings between males and females, with males having wing shapes that are more efficient in gliding flight performance.
↑But these estimates remain contentious, and at least 5 more species can be recognized based on the multispecies coalescent model as suggested by Matos-Maraví et al. (2019)
↑Markku Savela's Pierella page says this is invalid and the correct citation is Herrich-Schäffer, 1865, however no source is given for this conclusion and the LepIndex (among other sources) consider Westwood, [1851] to be correct.
Lamas, G. (ed), 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea – Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.
Matos-Maraví, P., Wahlberg, N., Antonelli, A. & Penz, C.M., 2019. Species limits in butterflies (Lepidoptera: Nymphalidae): reconciling classical taxonomy with the multispecies coalescent. Systematic Entomology, in press. DOI: http://doi.org/10.1111/syen.12352.