Heptophyllini

Heptophyllini
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Scarabaeiformia
Family:	Scarabaeidae
Subfamily:	Melolonthinae
Tribe:	Heptophyllini; Medvedev, 1951

Heptophyllini is a tribe of scarab beetles in the family Scarabaeidae.

Taxonomy

The tribe Heptophyllini is one of the insufficiently known and also poorly defined groups of the Old World Melolonthinae. The tribe was proposed by Medvedev to accommodate four genera (Heptophylla, Hexataenius, Hilyotrogus and Toxospathius). Medvedev argued out that Heptophyllini occupies a certain intermediate position between the tribes Melolonthini and Rhizotrogini. However, this opinion was weakly supported by morphological data. Kalinina examined the larval stages of Hilyotrogus bicoloreus in detail and found no differences from the larvae of Rhizotrogini, but retained Heptophyllini as a valid tribe. Although weakly supported, the tribe Heptophyllini has been considered valid in the last 50 years. Two additional genera (Laotrichia and Taiwanotrichia), were described, and the total number of taxa assigned to Heptophyllini now exceeds 50.

Genera

The following genera are recognised in the tribe Heptophyllini:

Heptophylla Motschulsky, 1857
Hexataenius Fairmaire, 1891
Hilyotrogus Fairmaire, 1886
Laotrichia Keith, 2007
Taiwanotrichia Kobayashi, 1990
Toxospathius Fairmaire, 1878

References

↑ Schoolmeesters, P. (2025). "Heptophyllini at Catalogue of Life". World Scarabaeidae Database (version 2025-10-07). In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, T. Pape, D. Hobern, S. Garnett, H. Little, R. E. DeWalt, J. Miller, T. Orrell, & R. Aalbu, Catalogue of Life (2025-10-10 XR). Amsterdam, Netherlands: Catalogue of Life Foundation. Retrieved May 23, 2026.
↑ Bezděk, A.; Král, D.; Luo, Y. P.; Lu, Y. Y. (2025). "Heptophylla gongshana Bezděk & Král, n. sp. from China (Yunnan) and a Checklist of the Tribe Heptophyllini (Scarabaeidae: Melolonthinae)". Taxonomy. 5 (52): 1-20. Retrieved May 23, 2026. This article incorporates text available under the CC BY 4.0 license.

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[:1-1] Schoolmeesters, P. (2025). "Heptophyllini at Catalogue of Life". World Scarabaeidae Database (version 2025-10-07). In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, T. Pape, D. Hobern, S. Garnett, H. Little, R. E. DeWalt, J. Miller, T. Orrell, & R. Aalbu, Catalogue of Life (2025-10-10 XR). Amsterdam, Netherlands: Catalogue of Life Foundation. Retrieved May 23, 2026.

[:2-2] Bezděk, A.; Král, D.; Luo, Y. P.; Lu, Y. Y. (2025). "Heptophylla gongshana Bezděk & Král, n. sp. from China (Yunnan) and a Checklist of the Tribe Heptophyllini (Scarabaeidae: Melolonthinae)". Taxonomy. 5 (52): 1-20. Retrieved May 23, 2026. This article incorporates text available under the CC BY 4.0 license.