Khasurty locality

Khasurty locality
Stratigraphic range: Neocomian
~145–130 Ma
Unit ofGusinoe Ozero Group
Location
RegionZakamensky District
CountryRussia
Type section
Named forKhasurty River

The Khasurty locality is an Early Cretaceous (Neocomian) lagerstätte in Russia, within the Sangin Formation. This lagerstätte is located outside the village of Tsakir in the region of Zakamensk. Over 6,000 insect specimens have been found from the site, belonging to around 130 families. Its insect biota resembles both Jurassic and Cretaceous ones, most heavily the Jehol biota of China.

Paleobiota

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon

Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Non-Insecta

Non-insect organisms
Genus Species Higher taxon Notes Images
Palaeoeorak P. scherbakovi Prochydorusidae (Anomopoda) Only named crustacean from Khasurty
The extant anomopod Daphnia magna
Bryokhutuliinia B. obtusifolia Hookeriales? Indeterminate family
Khasurtya K. ginkgoides Marchantiidae Specific name comes from the similarity in leaf shape to the ginkgo
Khasurtythallus K. monosolenioides Marchantiidae Likely an extinct clade of liverworts
Palaeodichelyma P. sinitzae Hypnanae (pleurocarpous mosses) Also known from the Jurassic of Russia
Tricostium T. longifolium Bryophyta incertae sedis Similar to a species found ~2000 km to the east

Insects

Palaeopterans

Palaeopterans
Genus Species Higher taxon Notes Images
Baikalogenites B. firmus Hexagenitidae (Ephemeroptera) Bears a longer seventh gill pair
Epiproctophora spp. Unapplicable Odonata Known from several unnamed species, most seemingly within Isophlebiidae or Campterophlebiidae
Isophlebia aspasia specimen from the Jurassic of Germany
Furvoneta F. khasurtensis Mesonetidae (Ephemeroptera) Largest member of the genus
Mesobaetis M. crispa Siphlonuridae Similar to the species M. amplectus, but has a larger rib on its gill plates
Mesogenesia M. sp Mickoleitiidae Only known from a poorly preserved specimen
Larva of the closely related Mickoleitia
Proameletus P. branchiatus Siphlonuridae Bears larger gills than the type species

Polyneopterans

Polyneopterans
Genus Species Higher taxon Notes Images
Aboilus? A?. khasurty Prophalangopsidae Differs from Aboilus in wing venation, but still fairly close to it
Pycnophlebia fossil from Jurassic Germany
Archisusumania A. contacta Susumanioidea Differs from other genera in wing venation
Three specimens of the closely related Cretophasmomima from China
Dimoula D. khasurtensis Nemouridae One of the few extinct nemourids known
Nemoura cinerea, an extant nemourid
Jurataenionema J. rohdendorfi Taeniopterygidae Known from a complete female specimen
Taeniopteryx nebulosa, an extant taeniopterygid
Mongoloxya M. fossor Tridactylidae Differs from the other species in the genus by having digging-adapted forelimbs
An unidentified extant tridactylid from India
Rasnitsyrina R. culonga, R. desiliens Perlariopseidae (Plecoptera) Most abundant stonefly from the site
Shurabia S. creta Geinitziidae Latest known winged notopteran besides those known from Burmese amber
Siberoperla S. bashkuevi Peltoperlidae Oldest peltoperlid, alongside the first fossil of a nymph from the family
An indeterminate peltoperlid nymph

Paraneopterans

Paraneopterans
Genus Species Higher taxon Notes Images
Buryatocera B. beta Kobdocoridae (Pentatomoidea) Has relatively short antennae
Creocanadaphis C. hirtus Canadaphididae Abdomen bears numerous small spines on sclerites
Extralapis E. carens, E. breviscutum Kobdocoridae Intermediate between the two other kobdocorid genera from Khasurty in pronotum (top-front section of thorax) proportions
Khasuaphis K. serotinus Juraphididae Bears longer antennae than other members of its family
Khasoris K. yuripopovi Kobdocoridae (Pentatomoidea) Bears a very long ovipositor
Liadopsylla L. (Basicella) lauteri, L. (Basicella) loginovae Liadopsyllidae Type genus of the family
Occidoscelis O. glaber Anthocorinae One of the only fossil anthocorids known
Anthocoris nemorum, an extant anthocorine bug
Reticycla R. drospoulosi Hylicellidae (Cicadomorpha) Has a smaller ovipositor and a different foreleg structure to most other hylicellids
Stigmapsylla S. klimaszewskii Liadopsyllidae Bears a dark pterostigma
Holotype of Amecephala pusilla, a different liadopsyllid from Burmese amber
Tobleronothrips T. deparis Aeolothripidae Named for Toblerone chocolate, which "noticeably helps when identifying complex imprints of thrips"
The modern Erythrothrips, another aeolothripid species

Holometabola

Hymenopterans
Hymenopterans
Genus Species Higher taxon Notes Images
Amplicella A. mininae, A. bashkuyevi, A. shcherbakovi, A. minor Tanychorinae (Ichneumonidae) Has a dark body with lighter legs/wing veins
Tanychora petiolata fossil from the Zaza Formation
Ghilarella G. kopylovi Sepulcidae Originally placed within G. mercurialis, as it only differed from the type of that species in wing colouration
Hymenoptera spp. Unapplicable Holometabola Includes numerous undescribed species from within various families like Angarosphecidae, Megalyridae and Proctotrupidae
Khasips K. alisecta, K. sculpta, K. kovalevi Archaeocynipidae (Cynipoidea) K. kovalevi has clubbed antennae, K. sculpta has an unusual pattern on its thorax
Khasurtella K. buryatica Tanychorinae (Ichneumonidae) Genus also contains a former Tanychora species
Micramphilius M. mirabilipennis Sepulcidae First member of its genus which preserves the hind wings
Onokhoius O. venustus Sepulcidae Bears a dark band on its pterostigma
Pamparaphilius P. khasurtensis Sepulcidae Holotype wing has numerous unusual veins, likely as a result of mutation
Parasyntexis P. khasurtensis Anaxyelidae
Praeichneumon P. dzhidensis, P. khamardabanicus, P. zakhaaminicus Praeichneumonidae Known from several mostly complete specimens
Sclerostigma S. trimaculata Dolichostigmatinae (Anaxyelidae) Species name means "three-spotted", as its abdominal tergites have patterns of three spots
Trematothorax T. zhangi, T. brachyurus, T. extravenosus Sepulcidae Another distinct species is known, however the material is too poor to name a new taxon
Neuropteridans
Neuropteridans
Genus Species Higher taxon Notes Images
Aberrantochrysa A. buryatica, A. pulchella Chrysopidae
Cretohondelagia C. viridis Priscaenigmatidae (Raphidioptera) Youngest priscaenigmatid fossil known
Dzhidosmylus D. solus Osmylidae Uncertain subfamily
Khasurtoberotha K. bellissima Berothidae Similar to the modern Protobiella
A specimen of the extant berothid Protobiella
Mesypochrysa M. cannabina, M. naranica Chrysopidae
Osmylochrysa O. navasia Osmylidae Previously only known from England, resembles chrysopid lacewings
Zakamnosmylus Z. elongatus Osmylidae Similar to the Jurassic Ensiosmylus
Coleopterans
Coleopterans
Genus Species Higher taxon Notes Images
Coleoptera spp. Unapplicable Coleoptera Contains many undescribed species in families such as Staphylinidae and Scarabaeidae
Coptoclava C. longipoda Coptoclavidae May not actually belong to the species
Coptoclava larva from the Jinju Formation
Distenorrhinus D. (Buryatnemonyx) ovatus, D. (Buryatnemonyx) gratshevi Nemonychidae Another undescribed species bears very large eyes
Gracilicupes G. minimus Cupedidae Much smaller than other species of the genus
Jurodes J. shef Jurodidae Similar to Jurodes from Daohugou
Fossil of Jurodes
Notocupes N. khasurtyiensis Archostemata incertae sedis Has larger eyes and a shorter first abdominal segment than other species in the genus
Several Notocupes specimens from Jurassic China alongside mid-Cretaceous Burmese amber
Palaeonecrophilus P. buryaticus Agyrtidae Earliest known agyrtid beetle
The extant agyrtid Necrophilus subterraneus
Ponohydrochus P. buryaticus Hydrochidae First fossil hydrochid genus
An extant Hydrochus beetle
Unda U. popovi Trachypachidae Also known from the Jurassic
The extant trachypachid Trachypachus gibbsii
Mecopterans
Mecopterans
Genus Species Higher taxon Notes Images
Antiquanabittacus A. dispar, A. robustus, A. saltator Bittacidae Similar to extant Orobittacus
The extant bittacid Bittacus selysi
Itaphlebia I. spp Nannochoristidae Known from very well-preserved material, however the difficult diagnosis of Itaphlebia species means the Khasurty species have not been named
The extant nannochoristid Nannochorista philpotti
Lichnomesopsyche L. sp Mesopsychidae Only known from one wing
Complete specimen of L. daohugouensis from Jurassic China
Meropeidae spp. Unapplicable Mecoptera Known from two different species each with a complete specimen, unusual for meropeids (which rarely fly)
The extant meropeid Merope tuber
Mesochorista M. sp Permochoristidae (related to Nannochoristidae) Formerly known as "Yanorthophlebia", which is a synonym of "Liassochorista" and in turn Mesochorista
Dipterans
Dipterans
Genus Species Higher taxon Notes Images
Amorimyia A. robusta Anisopodidae Differs from related genera in its hairy head and larger size
Collessomma C. sibirica, C. gnoma Perissommatidae Unlike modern perissommatids, this genus has fused (holoptic) eyes
The extant perissommatid Perissomma mcalpinei
Eoptychoptera E. fasbenderi Ptychopteridae Distinguished from other species in its genus by an additional cross-vein on the wing
The extant ptychopterid Ptychoptera contaminata
Gilkania G. transbaikalica Podonominae (Chironomidae) Only known from pupal exuvia
Juraxymyia J. krzeminskii Axymyiidae Also known from Daohugou
The extant axymyiid Axymyia furcata
Kaluginamyia K. enigmatica Kaluginamyiidae (Chironomoidea) Placed in its own family
Kovalevimyia K. sp Simuliidae One of the earliest hematophagous fly fossils, likely fed on birds
Mangas M. brevisubcosta, M. kovalevi Bolitophilidae Also known from Mongolia
The extant bolitophilid Bolitophila cinerea
Praearchitipula P. kaluginae, P. podenasi, P. ribeiroi Pediciidae Relatively large for pediciids
The extant pediciid Pedicia margarita
Protanyderus? P. madrizi Tanyderidae May not belong to the genus
Sinaxymyia S. szadziewskii Axymyiidae Also known from Daohugou
Zhiganka Z. longialata Ptychopteridae Known from complete specimens
The extant ptychopterid Bittacomorpha clavipes
Amphiesmenopterans
Amphiesmenopterans
Genus Species Higher taxon Notes Images
Axiomaldia A. flinti Trichoptera Uncertain family
Buranima B. gorhon Psychomyiidae Differs from other genera in having a maxillary palp without a longer second segment
The extant psychomyiid Tinodes waeneri
Burimodus B. novus Vitimotauliidae (Trichoptera)
Buryatocentrus B. suspiciosus Yantarocentridae (Limnephiloidea) Has two-segmented gonopods
Cretocrania C. stekolnikovi, C. glossancestralis, C. inconcessa Cretocraniinae (Eriocraniidae) Differs from modern eriocraniids in having round wing tips and very long galeae
The extant eriocraniid Eriocrania semipurpurella
Dauroglos D. hohlomak Glossosomatidae Similar to the genus Dajella
The extant Glossosoma intermedium
Duamodus D. anichkovi, D. necessarius Vitimotauliidae (Integripalpia) Similar to Multimodus
Dysoneura D. subbota Dysoneuridae Specific name derives from the Russian word for "Saturday"
Khasurtia K. kopylovi, K. alexeii, K. lukashevichae, K. novissima Dysoneuridae A fairly diverse caddisfly genus, almost entirely known from Khasurty
Juraphilopotamus J. callidus Philopotamidae Has small dark spots on forewing
The extant philopotamid Philopotamus montanus
Meloclada M. diuturna, M. frequentatoria Cladochoristidae (Trichoptera) Similar to the related Cladochoristella
Mesocolepus M. deflectum Ptilocolepidae (Hydroptiloidea) Bears large setal warts, with their shape used to distinguish the genus
Multimodus M. amplus Vitimotauliidae (Trichoptera) Largest species of the genus
Palaeosiberomiya P. zugzwanga Dysoneuridae Bears longitudinal stripes of dark scales on its wings
Protobaikalopsyche P. rossica Protobaikalopsychidae (Hydropsychoidea) Likely basal to a clade including Polycentropodidae
Protorodinia P. khasurtensis Leptoceridae Bears a large mesoscutum and unusual abdominal tip
The extant leptocerid Leptocerus tineiformis
Protosiberopteryx P. antitheton, P. equesaeneus, P. praevernalis Eolepidopterigidae Known from one male and two female specimens, similar to Undopterix
Purbimodus P. parvulus, P. khramovi Vitimotauliidae (Trichoptera) Redescribed in a 2021 paper
Sententimiya S. wichardi Psychomyiidae Bears a plate on the fifth sternite similar to those on psychomyiids
Shadareniskia S. inambularia Hydrobiosidae Known from a female specimen, as shown by the genitalia and a suture on the sixth sternite
The (male) holotype of the extant Hydrobiosis falcis
Siberocretopetus S. ekaterinae Glossosomatidae Specific name honours Ekaterina Sidorchuk, a palaeoacarologist (researcher who studies fossil mites) who died in 2019
The extant agapetine glossosomatid Agapetus fuscipes
Siberoprostoria S. archaeoscriptor Hydrobiosidae Known from a female specimen, as shown by the genitalia and a suture on the sixth sternite
Styxowia S. predponimania Polycentropodidae Has large, spiny gonopods
The extant polycentropodid Polycentropus flavomaculatus
Susurimiya S. transbaikalica Psychomyiidae Bears a swollen abdomen tip like modern psychomyiids, but differs in the structure of its maxillary palps
Terminoptysma T. contradictoria Terminoptysmatidae (Trichoptera) Placed in its own monotypic family
Terrindusia T. khasurtica, T. buriatica Trichoptera Known only from larval casings

References

  1. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 Kopylov, D. S.; Rasnitsyn, A. P.; Aristov, D. S.; Bashkuev, A. S.; Bazhenova, N. V.; Dmitriev, V. Yu.; Gorochov, A. V.; Ignatov, M. S.; Ivanov, V. D.; Khramov, A. V.; Legalov, A. A.; Lukashevich, E. D.; Mamontov, Yu. S.; Melnitsky, S. I.; Ogłaza, B.; Ponomarenko, A. G.; Prokin, A. A.; Ryzhkova, O. V.; Shmakov, A. S.; Sinitshenkova, N. D.; Solodovnikov, A. Yu.; Strelnikova, O. D.; Sukacheva, I. D.; Uliakhin, A. V.; Vasilenko, D. V.; Wegierek, P.; Yan, E. V.; Zmarzły, M. (December 2020). "The Khasurty Fossil Insect Lagerstätte". Paleontological Journal. 54 (11): 1221–1394. Bibcode:2020PalJ...54.1221K. doi:10.1134/S0031030120110027.
  2. ↑ Kotov, Alexey A. (February 2009). "A revision of the extinct Mesozoic family Prochydoridae Smirnov, 1992 (Crustacea: Cladocera) with a discussion of its phylogenetic position". Zoological Journal of the Linnean Society. 155 (2): 253–265. doi:10.1111/j.1096-3642.2008.00412.x.
  3. 1 2 3 Ignatov, Michael S.; Schherbakov, Dmitry E. (December 2011). "Lower Cretaceous mosses from Khasurty (Baikal Area of South Siberia)". Arctoa. 20 (1): 19–42. doi:10.15298/arctoa.20.02.
  4. ↑ Mamontov, Yuriy S.; Ignatov, Michael S. (July 2019). "How to rely on the unreliable: Examples from Mesozoic bryophytes of Transbaikalia". Journal of Systematics and Evolution. 57 (4): 339–360. doi:10.1111/jse.12483.
  5. 1 2 3 4 5 Sinitshenkova, N. D. (May 2017). "New mayflies (Insecta: Ephemerida = Ephemeroptera) from the Upper Mesozoic locality of Khasurty, Western Transbaikalia". Paleontological Journal. 51 (3): 280–287. Bibcode:2017PalJ...51..280S. doi:10.1134/S003103011703011X.
  6. 1 2 3 Sinitshenkova, N. D. (September 2011). "New stoneflies (Insecta: Perlida = Plecoptera) from the late Mesozoic locality Khasurty, Transbaikalia". Paleontological Journal. 45 (5): 552–558. doi:10.1134/S0031030111050133.
  7. ↑ Sinitshenkova, Nina; Yan, Evgeny (15 October 2024). "The most ancient roachfly (Insecta: Plecoptera, Peltoperlidae) from the Lower Cretaceous of the West Transbaikalia, Russia". Zootaxa. 5523 (2): 284–290. doi:10.11646/zootaxa.5523.2.9.
  8. 1 2 3 Ryzhkova, Olga V.; Yao, Yunzhi; Kopylov, Dmitry S. (July 2023). "New fossil Kobdocoridae (Heteroptera: Pentatomomorpha) from the Lower Cretaceous of Transbaikalia, Eastern Russia". Cretaceous Research. 147 105509. doi:10.1016/j.cretres.2023.105509.
  9. 1 2 3 Shcherbakov, D. E. (December 2020). "New Homoptera from the Early Cretaceous of Buryatia with notes on the insect fauna of Khasurty". Russian Entomological Journal. 29 (1): 127–138. doi:10.15298/rusentj.29.2.02.
  10. ↑ Kopylov, D. S. (May 2018). "New Ichneumonids (Hymenoptera: Ichneumonidae) from the Lower Cretaceous Khasurty Locality". Paleontological Journal. 52 (3): 284–289. doi:10.1134/S003103011803005X.
  11. 1 2 Kopylov, D. S. (July 2011). "Ichneumon wasps of the Khasurty locality in Transbaikalia (Insecta, Hymenoptera, Ichneumonidae)". Paleontological Journal. 45 (4): 406–413. doi:10.1134/S0031030111040058.
  12. ↑ Li, Yi; Wang, Mei; Rasnitsyn, Alexandr P.; Shih, Chungkun; Zhuang, Jialiang; Ren, Dong (July 2024). "Two new species of Ghilarellinae (Hymenoptera, Cephoidea, Sepulcidae) from the Lower Cretaceous". Cretaceous Research. 159 105875. doi:10.1016/j.cretres.2024.105875.
  13. 1 2 Kopylov, D. S.; Rasnitsyn, A. P. (May 2017). "New sepulcids (Hymenoptera: Sepulcidae) from the Lower Cretaceous of Asia: II. Ghilarellinae and Trematothoracinae". Paleontological Journal. 51 (3): 291–303. doi:10.1134/S0031030117030029.
  14. ↑ Kopylov, D. S. (February 2014). "New archaeocynipids (Hymenoptera, Cynipoidea, Archaeocynipidae) from the Lower Cretaceous of Transbaikalia". Paleontological Journal. 48 (1): 58–64. doi:10.1134/S0031030114010067.
  15. 1 2 3 Kopylov, D. S.; Rasnitsyn, A. P. (January 2017). "New sepulcids (Hymenoptera: Sepulcidae) from the Lower Cretaceous of Asia. I. Parapamphiliinae and Xyelulinae". Paleontological Journal. 51 (1): 69–76. doi:10.1134/S0031030117010087.
  16. ↑ Kopylov, Dmitry S. (9 May 2019). "New anaxyelids (Hymenoptera: Anaxyelidae) from the Cretaceous of Asia". Zootaxa. 4603 (2). doi:10.11646/ZOOTAXA.4603.2.7.
  17. ↑ Kopylov, D. S. (January 2012). "New species of Praeichneumonidae (Hymenoptera, Ichneumonoidea) from the Lower Cretaceous of Transbaikalia". Paleontological Journal. 46 (1): 66–72. doi:10.1134/S0031030112010078.
  18. 1 2 Khramov, Alexander V. (June 2016). "A new assemblage of Early Cretaceous green lacewings (Chrysopidae: Neuroptera) from Transbaikalia". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 107 (2–3): 195–202. doi:10.1017/S1755691017000342.
  19. ↑ Khramov, Alexander V. (3 October 2021). "The youngest record of Priscaenigmatidae (Insecta: Neuropterida) from the Mesozoic of Kazakhstan and Russia". Historical Biology. 33 (10): 2432–2437. doi:10.1080/08912963.2020.1800683.
  20. 1 2 Strelnikova, O. D. (May 2019). "New Cupedids (Insecta: Coleoptera, Cupedidae) from the Lower Cretaceous of Buryatia". Paleontological Journal. 53 (3): 292–299. doi:10.1134/S0031030119030146.
  21. ↑ Yan, Evgeny V.; Strelnikova, Olesya D.; Ponomarenko, Alexander G. (14 December 2021). "Jurodidae (Coleoptera: Archostemata) from Transbaikalia: new findings and redescription of type material". Palaeoentomology. 4 (6). doi:10.11646/palaeoentomology.4.6.9.
  22. ↑ Strelnikova, O. D.; Yan, E. V.; Vasilenko, D. V. (November 2020). "A New Agyrtid Beetle (Coleoptera, Agyrtidae) from the Lower Cretaceous Khasurty Locality". Paleontological Journal. 54 (6): 627–631. doi:10.1134/S003103012006009X.
  23. ↑ Prokin, Alexander A.; Strelnikova, Olesya D. (July 2021). "The first remarkable fossil Hydrochidae (Coleoptera) from the Lower Cretaceous (Hauterivian) of the Buryatia Republic, Russia". Cretaceous Research. 123 104795. doi:10.1016/j.cretres.2021.104795.
  24. ↑ Ponomarenko, A. G. (April 2023). "A New Species of Ground Beetle (Insecta, Coleoptera, Trachypachidae) from the Lower Cretaceous Khasurty Locality, Western Transbaikalia". Paleontological Journal. 57 (2): 196–198. doi:10.1134/S0031030123010112.
  25. ↑ Bashkuev, Alexey S. (August 2023). "Morphology and relationships of the bittacid genus Antiquanabittacus Petrulevičius et Jarzembowski, 2004 (Mecoptera), with description of three new species from the Lower Cretaceous of south-east Siberia". Cretaceous Research. 148 105549. doi:10.1016/j.cretres.2023.105549.
  26. ↑ Ansorge, Jörg (1995). "Revision der liassischen "Permochoristidae" (Stammgruppenvertreter der Antliophora = Mecoptera + Diptera)" (PDF). Studia dipterologica. 2 (1): 21–26.
  27. 1 2 3 4 5 Lukashevich, E. D. (November 2020). "Rare Nematoceran Dipterans (Insecta: Diptera) from the Khasurty Locality, Transbaikalia". Paleontological Journal. 54 (6): 600–612. doi:10.1134/S0031030120060040.
  28. ↑ Lukashevich, Elena D.; Blagoderov, Vladimir A. (8 January 2020). "Review of Mesozoic Perissommatidae (Insecta: Diptera)". Zootaxa. 4718 (4). doi:10.11646/zootaxa.4718.4.3.
  29. 1 2 Blagoderov, Vladimir A.; Lukashevich, Elena D. (1 December 2013). "New Axymyiidae (Insecta: Diptera) from the Mesozoic of East Siberia". Polish Journal of Entomology / Polskie Pismo Entomologiczne. 82 (4): 257–271. doi:10.2478/v10200-012-0040-9.
  30. ↑ Lukashevich, Elena D.; Pepinelli, Mateus; Currie, Douglas C. (1 February 2020). "A new family and genus of Mesozoic Simulioidea (Insecta: Diptera)". Journal of Systematic Palaeontology. 18 (3): 213–231. doi:10.1080/14772019.2019.1588796.
  31. ↑ Greenwalt, Dale E.; Blagoderov, Vladimir A. (18 March 2019). "Review of the fossil record of Bolitophilidae, with description of new taxa and discussion of position of Mangas Kovalev (Diptera: Sciaroidea)". Zootaxa. 4567 (3). doi:10.11646/zootaxa.4567.3.6.
  32. ↑ Lukashevich, Elena D. (29 August 2019). "Review of the fossil record of Bittacomorphinae (Diptera: Ptychopteridae)". Zootaxa. 4661 (3). doi:10.11646/zootaxa.4661.3.9.
  33. 1 2 3 Sukatsheva, I. D.; Aristov, D. S. (July 2020). "New Caddisflies (Insecta, Trichoptera) of the Families Vitimotauliidae and Philopotamidae from the Khasurty Locality (Lower Cretaceous, Russia) with a Brief Review of the World Fossil Fauna". Paleontological Journal. 54 (4): 375–384. doi:10.1134/S0031030120040139.
  34. 1 2 Sukatsheva, I. D.; Vasilenko, D. V. (September 2019). "New Caddisflies of the Family Dysoneuridae (Insecta: Trichoptera) and Larval Cases (Incertae Familiae) from the Lower Cretaceous of Transbaikalia". Paleontological Journal. 53 (5): 499–505. doi:10.1134/S0031030119050125.
  35. ↑ Sukatsheva, I. D.; Aristov, D. S. (July 2021). "New Data on Cretaceous Caddisflies Family Vitimotauliidae (Insecta, Trichoptera) from Buryatia (Khasurty Locality)". Paleontological Journal. 55 (4): 391–395. doi:10.1134/S0031030121330017.