Manulea

Manulea
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Lepidoptera
Superfamily:	Noctuoidea
Family:	Erebidae
Subfamily:	Arctiinae
Subtribe:	Lithosiina
Genus:	Manulea; Wallengren, 1863
Synonyms
	Agenjoa Dubatolov & Zolotuhin, 2011 (= Nyea Agenjo, 1983, syn. nov.); Colinia Agenjo, 1977 (preocc. Nuttall 1832);

Manulea is a genus of moths in the family Erebidae erected by Hans Daniel Johan Wallengren in 1863. The type species is Lithosia gilveola Ochsenheimer, 1810 (= palliatella Scopoli, 1763), designated by subsequent designation by Moore in 1878.

Like other members of the subtribe Lithosiina, larvae of species in Manulea feed primarily on lichens. The larvae of Manulea complana, for example, feed on various species of lichen (Parmelia spp.) as well as dry and tender leaves of various plants including Quercus, Fagus, Prunus, Salix, and Rubus.

Species

Subgenus Manulea Wallengren, 1863
complana species group

Manulea complana (Linnaeus, 1758)
Manulea costalis (Zeller, 1847) (=morosina Herrich-Schäffer, [1847])
Manulea palliatella (Scopoli, 1763) (=unita [Denis & Schiffermüller], 1775)
Manulea pseudocomplana (Daniel, 1939)

lutarella species group

Manulea lutarella (Linnaeus, 1758)
Manulea flavociliata (Lederer, 1853)

minor species group

Manulea minor (Okano, 1955)
Manulea pseudofumidisca Dubatolov & Zolotuhin, 2011

pygmaeola species group

Manulea affineola (Bremer, 1864)
Manulea fuscodorsalis (Matsumura, 1930)
Manulea japonica (Leech, [1889])
Manulea kansuensis (Hering, 1936)
Manulea minima (Daniel, 1954)
Manulea nankingica (Daniel, 1954)
Manulea omelkoi Dubatolov & Zolotuhin, 2011
Manulea predotae (Schawerda, 1927)
Manulea pygmaeola (Doubleday, 1847)
Manulea ussurica (Daniel, 1954)
Manulea uniformeola (Daniel, 1954)
Manulea wiltshirei (Tams, 1939)

Subgenus Agenjoa Dubatolov & Zolotuhin, 2011

Manulea hunanica (Daniel, 1954)
Manulea lurideola (Zincken, 1817)

A subgenus Churingosia Volynkin & Černý was described in 2022, with the type species Ilema nigripes Hampson, 1900. It included species from the Oriental realm, among them Manulea (Churingosia) chrysophleps (Hampson, 1895) and Manulea (Churingosia) nigripes (Hampson, 1900).

Four new species were described at the same time: Manulea (Churingosia) fardyftera from Northeast India, M. (C.) reducta from northern Myanmar and Northeast India, M. (C.) mikrotera from northern Myanmar and Northeast India, and M. (C.) mavropoda from northern Indochina.

Incorrectly placed

Manulea replana (Lewin, 1805) (now Brunia replana)

External links

Dubatolov, V. V. & Zolotuhin, V. V. (2011). "Does Eilema Hübner, [1819] (Lepidoptera, Arctiidae, Lithosiinae) present one or several genera?" (PDF). Euroasian Entomological Journal. 10 (3): 367–379, 380, VII.
Dubatolov, Vladimir V.; Kishida, Yasunori & Wang, Min (May 2012). "New records of lichen-moths from the Nanling Mts., Guangdong, South China, with descriptions of new genera and species (Lepidoptera, Arctiidae: Lithosiinae)" (PDF). Tinea. 22 (1): 25–52. S2CID 86037229. Archived from the original (PDF) on 2019-10-09.
Pitkin, Brian & Jenkins, Paul (2023). "Search results Family: Arctiidae". Butterflies and Moths of the World. Natural History Museum, London. doi:10.5519/s93616qw.
Yu, Dicky Sick Ki. "Manulea Wallengren 1863". Home of Ichneumonoidea. Taxapad. Archived from the original on March 5, 2016.

References

↑ Volynkin, A.V. (2021) Tortrilema, a new subgenus of the genus Manulea Wallengren with description of a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 48, 9–17. https://doi.org/10.37828/em.2021.48.2
↑ Volynkin, Anton V.; Saldaitis, Aidas (2021-06-24). "Two new species of Manulea Wallengren, 1863 from southwestern China and northern Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini)". Zootaxa. 4991 (2): 389–397. doi:10.11646/zootaxa.4991.2.12. ISSN 1175-5334.
↑ "We'll forward you in a second". treatment.plazi.org. Retrieved 2026-04-04.
1 2 Volynkin, Anton V.; Černý, Karel (2022-11-07). "Churingosia, a new subgenus of the genus Manulea Wallengren with descriptions of four new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae)". Zootaxa. 5205 (2): 162–176. doi:10.11646/zootaxa.5205.2.4. ISSN 1175-5334.

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[1] Volynkin, A.V. (2021) Tortrilema, a new subgenus of the genus Manulea Wallengren with description of a new species from Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 48, 9–17. https://doi.org/10.37828/em.2021.48.2

[2] Volynkin, Anton V.; Saldaitis, Aidas (2021-06-24). "Two new species of Manulea Wallengren, 1863 from southwestern China and northern Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini)". Zootaxa. 4991 (2): 389–397. doi:10.11646/zootaxa.4991.2.12. ISSN 1175-5334.

[3] "We'll forward you in a second". treatment.plazi.org. Retrieved 2026-04-04.

[:0-4] 1 2 Volynkin, Anton V.; Černý, Karel (2022-11-07). "Churingosia, a new subgenus of the genus Manulea Wallengren with descriptions of four new species from the Oriental realm (Lepidoptera: Erebidae: Arctiinae)". Zootaxa. 5205 (2): 162–176. doi:10.11646/zootaxa.5205.2.4. ISSN 1175-5334.