Permosynidae

Permosynidae; Temporal range: 268.0 to 122.46 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	Holotype of Dinoharpalus coptoclavoides, photograph and illustration
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	†Protocoleoptera
Superfamily:	†Permosynoidea
Family:	†Permosynidae; Tillyard, 1924
Genera
	See text;

Permosynidae is an extinct family of beetles known from the Middle Permian to Early Cretaceous. It is sometimes treated as a form family, because all known members were described from fossils of isolated elytra with punctate striae.

Taxonomy

After:

Alveolacupes Lin, 1983 (also spelled Alveolicupes)
- Alveolacupes primus Lin, 1983 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
- Alveolacupes secundes Lin, 1983 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
Artematopodites Ponomarenko, 1990
- Artematopodites crispulus Yan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Artematopodites insculptus (Zhang, 1997) – Sangonghe Formation, China, Early Jurassic (Toarcian); Xishanyao Formation, China, Middle Jurassic (Aalenian/Bajocian)
- Artematopodites latissimus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Artematopodites latus Ponomarenko, 1990 – Byankino Formation, Russia, Late Jurassic (Tithonian)
- Artematopodites lepidus Yan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Artematopodites leskoviensis Yan in Yan and Zhang, 2010 – Leskovskaya Formation, Russia, Early Cretaceous (Barremian)
- Artematopodites longus (Hong, 1983) – Jiulongshan Formation, China, Middle-Late Jurassic (Callovian/Oxfordian)
- Artematopodites lozowskii Ponomarenko, 2015 – Vokhma Formation, Russia, Late Permian (Changhsingian)
- Artematopodites major Ponomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
- Artematopodites maximus Yan in Yan and Zhang, 2010 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Artematopodites prolixus (Zhang, 1997) – Badaowan Formation, China, Early Jurassic (Hettangian/Sinemurian)
- Artematopodites propinquus (Zhang, 1997) – Badaowan Formation, China, Early Jurassic (Hettangian/Sinemurian)
- Artematopodites shaanbeiensis Hong, 1995 – Huachi-Huanhe Formation, China, Early Cretaceous (Valanginian)
Bistrisyne Lin, 1986
- Bistrisyne tenua Lin, 1986 – Sanqiutang Formation, China, Late Triassic (Rhaetian)
Delpuentesyne Martins-Neto and Gallego, 2007
- Delpuentesyne menendezi Martins-Neto and Gallego, 2007 – Potrerillos Formation, Argentina, Middle Triassic (Ladinian)
Diarcuipenna Lin, 1986
- Diarcuipenna bennettitophila Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
- Diarcuipenna heterosa Lin, 1986 – Zijiachong Formation, China, Late Triassic (Norian)
Dinoharpalus Handlirsch, 1906
- Dinoharpalus coptoclavoides Ponomarenko, 2015 – Hassberge Formation, Germany, Late Triassic (Carnian)
- Dinoharpalus latus Ponomarenko, 2015 – Vokhma Formation, Russia, Late Permian (Changhsingian)
- Dinoharpalus liasinus (Giebel, 1856) – Cotham Formation, England, Late Triassic (Rhaetian)
- Dinoharpalus martynovi Ponomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
- Dinoharpalus rugosus Ponomarenko, 1985 – Osinovo Formation, Russia, Early-Middle Jurassic (Toarcian/Aalenian)
Dzeregia Ponomarenko, 1985
- Dzeregia ampla Ponomarenko, 1990 – Mangutskaya Formation, Russia, Early Cretaceous (Berriasian)
- Dzeregia byrrhoides Ponomarenko, 1990 – Ukurei Formation, Russia, Late Triassic (Lithonian)
- Dzeregia crassa Ponomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
- Dzeregia juxta (Lin, 1986) – Zijiachong Formation, China, Late Triassic (Norian)
- Dzeregia lata Ponomarenko, 1985 – Makarova Formation, Russia, Early Jurassic (Toarcian)
- Dzeregia longa Ponomarenko, 1985 – Tersyukskaya Formation, Russia, Middle Jurassic (Bajocian)
- Dzeregia pilula Ponomarenko, 1990 – Turga Formation, Russia, Early Cretaceous (Aptian)
- Dzeregia platis Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Dzeregia striata Ponomarenko, 1985 – Zhargalant Formation, Mongolia, Early Jurassic (Toarcian)
Hydrobiites Heer, 1865
- Hydrobiites anglicus Handlirsch, 1906 – Hasfield, England, Late Triassic (Rhaetian)
- Hydrobiites bellus (Geinitz, 1894) – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
- Hydrobiites convexus Ponomarenko, 1985 – Tersyukskaya Formation, Russia, Middle Jurassic (Bajocian)
- Hydrobiites crassus Ponomarenko, 1985 – Cheremkhovskaya Formation, Russia, Early Jurassic (Toarcian)
- Hydrobiites dobbertinensis Handlirsch, 1939 – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
- Hydrobiites giebeli Handlirsch, 1906 – Cotham Formation, England, Late Triassic (Rhaetian)
- Hydrobiites handlirschi Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
- Hydrobiites liasinus (Giebel, 1856) – Hasfield, England, Late Triassic (Rhaetian)
- Hydrobiites longus Ponomarenko, 1990 – Kutinskaya Formation, Russian, Early Cretaceous (Barremian)
- Hydrobiites minor Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Hydrobiites minutissimus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
- Hydrobiites mongolicus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Hydrobiites permianus Ponomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
- Hydrobiites punctatostriatus Handlirsch, 1939 – Dobbertin, Mecklenburg-Vorpommern, Germany, Early Jurassic (Toarcian)
- Hydrobiites purbeccensis (Giebel, 1856) – Lulworth Formation, England, Early Cretaceous (Berriasian)
- Hydrobiites sulcatus Ponomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Early Cretaceous (Aptian)
- Hydrobiites tillyardi Ponomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
- Hydrobiites veteranus (Heer, 1852) – Insektenmergel Formation, Switzerland, Early Jurassic (Hettangian)
- Hydrobiites vladimiri Ponomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
Ischichucasyne Martins-Neto and Gallego, 2005
- Ischichucasyne cladocosta Martins-Neto and Gallego, 2009 – Los Rastros Formation, Argentina, Late Triassic (Carnian)
- Ischichucasyne santajuanaensis Martins-Neto and Gallego, 2005 – Santa Juana Formation, Chile, Late Triassic (Carnian)
Palademosyne Rohdendorf, 1961
- Palademosyne martynovae Rohdendorf, 1961 – Kazankovo-Markinskaya Formation, Russia, Middle Permian (Wordian)
- Palademosyne natalensis Ponomarenko, 2005 – Estcourt Formation, South Africa, Late Permian (Changhsingian)
Permocrossos Martynov, 1932
- Permocrossos elongatus Martynov, 1932 – Iva-Gora Beds Formation, Russia, Middle Permian (Roadian)
Permosyne Tillyard, 1924
- Permosyne affinis Tillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
- Permosyne belmontensis Tillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
- Permosyne dentata Ponomarenko, 2003 – Salarevo Formation, Russia, Late Permian (Changhsingian)
- Permosyne elongata Ponomarenko, 2005 – Estcourt Formation, South Africa, Late Permian (Changhsingian)
- Permosyne mitchelli Tillyard, 1924 – Croudace Bay Formation, Australia, Late Permian (Changhsingian)
- Permosyne rasnitsyni Ponomarenko, 2011 – Vyazniki Formation, Russia, Late Permian (Changhsingian)
Platycrossos Dunstan, 1923
- Platycrossos caroli Ponomarenko, 2011 – Lunz Formation, Austria, Late Triassic (Carnian)
- Platycrossos elongatus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
- Platycrossos latus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Platycrossos ligulatus Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- Platycrossos longus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Platycrossos loxonicus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Platycrossos mongolicus Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Platycrossos ovum Ponomarenko, 2014 – Sharteg Formation, Mongolia, Late Jurassic (Tithonian)
- Platycrossos petalus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
- Platycrossos punctatus Ponomarenko, 1985 – Itat Formation, Russia, Middle Jurassic (Bajocian/Bathonian)
- Platycrossos subtumidus Dunstan, 1923 – Blackstone Formation, Australia, Late Triassic (Norian)
- Platycrossos tumidus (Tillyard, 1916) – Blackstone Formation, Australia, Late Triassic (Norian)
Pseudorhynchophora Handlirsch, 1906
- Pseudorhynchophora convexa Ponomarenko, 2008 – Nyadeita Formation, Russia, Early Triassic (Olenekian)
- Pseudorhynchophora olliffi Handlirsch, 1906 – Blackstone Formation, Australia, Late Triassic (Norian)
Sakmaracoleus Ponomarenko, 2013
- Sakmaracoleus orenburgensis Ponomarenko, 2013 – Vyazovka Formation, Russia, Middle Permian (Capitanian)
Tychtocoleus Rohdendorf, 1961
- Tychtocoleus europaeus (Ponomarenko, 2003) – Salarevo Formation, Russia, Late Permian (Changhsingian)
- Tychtocoleus neuburgae Rohdendorf, 1961 – Gramoteinskaya Formation, Russia, Late Permian (Wuchiapingian)
- Tychtocoleus popovi Ponomarenko, 2018 – Salarevo Formation, Russia, Late Permian (Changhsingian)
Umkomaasia Zeuner, 1960
- Umkomaasia depressa Zeuner, 1960 – Molteno Formation, South Africa, Late Triassic (Carnian)

References

1 2 "Permosynidae". Paleobiology Database. Retrieved 9 March 2023.
↑ Ponomarenko, A.G. (2003). "The First Beetles (Permosynidae, Coleoptera) from the Upper Tatarian of European Russia" (PDF). Paleontological Journal. 37 (2): 170–173.
↑ A. G. Ponomarenko, D. S. Aristov, A. S. Bashkuev, Yu. M. Gubin, A. V. Khramov, E. D. Lukashevich, Yu. A. Popov, L. N. Pritykina, S. M. Sinitsa, N. D. Sinitshenkova, I. D. Sukatsheva, D. V. Vassilenko, E. V. Yan (2014). "Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia". Paleontological Journal 48 (14): 1573–1682. doi:10.1134/S0031030114140160.
↑ A. G. Ponomarenko (2015). "New beetles (Insecta, Coleoptera) from the Nedubrovo locality, terminal Permian or basal Triassic of European Russia". Paleontological Journal 49 (1): 39–50. doi:10.1134/S0031030115010098.

[Fossilworks-1] 1 2 "Permosynidae". Paleobiology Database. Retrieved 9 March 2023.

[2] Ponomarenko, A.G. (2003). "The First Beetles (Permosynidae, Coleoptera) from the Upper Tatarian of European Russia" (PDF). Paleontological Journal. 37 (2): 170–173.

[3] A. G. Ponomarenko, D. S. Aristov, A. S. Bashkuev, Yu. M. Gubin, A. V. Khramov, E. D. Lukashevich, Yu. A. Popov, L. N. Pritykina, S. M. Sinitsa, N. D. Sinitshenkova, I. D. Sukatsheva, D. V. Vassilenko, E. V. Yan (2014). "Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia". Paleontological Journal 48 (14): 1573–1682. doi:10.1134/S0031030114140160.

[4] A. G. Ponomarenko (2015). "New beetles (Insecta, Coleoptera) from the Nedubrovo locality, terminal Permian or basal Triassic of European Russia". Paleontological Journal 49 (1): 39–50. doi:10.1134/S0031030115010098.

Permosynidae Temporal range: 268.0 to 122.46 Ma PreꞒ Ꞓ O S D C P T J K Pg N

Holotype of Dinoharpalus coptoclavoides, photograph and illustration
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	†Protocoleoptera
Superfamily:	†Permosynoidea
Family:	†Permosynidae Tillyard, 1924
Genera
See text