Brower, A.V.Z. et al. (2006). Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology31: 288-301.[dode link]
↑Hoewel Fox, R.M. (1956). A monograph of the Ithomiidae (Lepidoptera). Part 1. Bulletin of the American Museum of Natural History111: 1–76, pl. 71–79, Aeria in de Oleriini plaatste, is het aannemelijker dat dit geslacht thuishoort in Tithoreina (Ackery, P.R. (1988). Hostplants and classification: a review of nymphalid butterflies. Biological Journal of the Linnean Society33: 95–203; Brown Jr K.S. & Freitas, A.V.L. (1994). Juvenile stages of Ithomiinae: overview and systematics. Tropical Lepidoptera5: 9–20). Zie: Brower, A.V.Z. et al. (2006). Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology31: 298.[dode link]
Tithoreina op Wikispecies