Geomitridae

Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.

The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus. A free right ommatophoral retractor has been linked to adaptation to xeric habitats.

The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015). Genera with the family Geomitridae include:

Family Geomitridae

• Not belonging to a subfamily:
  • Keraea Gude, 1911
  • Peridotitea Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018

• Subfamily Geomitrinae
  • tribe Cochlicellini Schileyko, 1972
    • Cochlicella Férussac, 1821
    • Monilearia Mousson, 1872
    • Obelus W. Hartmann, 1842
    • Ripkeniella Hutterer & E. Gittenberger, 1998
  • tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)
    • Actinella R. T. Lowe, 1852
    • Callina R. T. Lowe, 1855
    • Caseolus R. T. Lowe, 1852
    • Discula R. T. Lowe, 1852
    • Disculella Pilsbry, 1895
    • Domunculifex Brozzo, De Mattia, Harl & Neiber, 2020
    • Geomitra Swainson, 1840
    • Helicomela R. T. Lowe, 1855
    • Heterostoma W. Hartmann, 1843: synonym of Steenbergia Mandahl-Barth, 1950 (invalid: junior homonym of Heterostoma de Filippi, 1837 [Platyhelminthes])
    • Hystricella R. T. Lowe, 1855
    • Lemniscia R. T. Lowe, 1855
    • Moreletina de Frias Martins, 2002
    • Plebecula R. T. Lowe, 1852
    • Pseudocampylaea L. Pfeiffer, 1877
    • Serratorotula Groh & Hemmen, 1986
    • Spirorbula R. T. Lowe, 1852
    • Steenbergia Mandahl-Barth, 1950
    • Testudodiscula Brozzo, De Mattia, Harl & Neiber, 2020
    • Wollastonaria De Mattia, Neiber & Groh, 2018
  • tribe Ponentinini Razkin et al., 2015
    • Ponentina P. Hesse, 1921

• subfamily Helicellinae
  • tribe Cernuellini Schileyko, 1991
    • Alteniella Clerx & Gittenberger, 1977
    • Cernuella Schlüter, 1838
    • Microxeromagna Ortiz de Zárate López, 1950
    • Xeroplana Monterosato, 1892
    • Xerosecta Monterosato, 1892
  • tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)
    • Backeljaia Chueca et al., 2018
    • Candidula Kobelt, 1871
    • Helicella Férussac, 1821
    • Orexana Chueca et al., 2018
    • Xerogyra Monterosato, 1892
    • Xeroleuca Kobelt, 1877
    • Xeroplexa Monterosato, 1892
    • Xerotricha Monterosato, 1892
    • Zarateana Chueca et al., 2018
  • tribe Helicopsini H. Nordsieck, 1987
    • Helicopsis Fitzinger, 1833
    • Pseudoxerophila Westerlund, 1879
    • Xerolenta Monterosato, 1892
    • Xeromunda Monterosato, 1892
    • Xeropicta Monterosato, 1892
  • tribe Plentuisini Razkin et al., 2015
    • Plentuisa Puente & Prieto, 1992
  • tribe Trochoideini H. Nordsieck, 1987
    • Trochoidea T. Brown, 1827
    • Xerocrassa Monterosato, 1892

• Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

• https://liferecovernatura.madeira.gov.pt
• Razkin, O., Gómez-Moliner, B. J., Prieto, C. E., Martínez-Ortí, A., Arrébola, J. R., Muñoz, B., Chueca, L. J. & Madeira, M. J. (2015). Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora). Molecular Phylogenetics and Evolution. 83: 99-117