List of macroscelids

IUCN Red List categories
Conservation status
EX	Extinct (0 species)
EW	Extinct in the wild (0 species)
CR	Critically endangered (0 species)
EN	Endangered (1 species)
VU	Vulnerable (1 species)
NT	Near threatened (0 species)
LC	Least concern (13 species)
Other categories
DD	Data deficient (4 species)
NE	Not evaluated (0 species)

Macroscelidea is an order of small mammals. Members of this order are called macroscelids, or commonly as sengis or elephant shrews. They are exclusively found in Africa, in a variety of biomes from forests to deserts. They range in size from the Etendeka round-eared sengi, at 8 cm (3 in) plus a 8 cm (3 in) tail, to the grey-faced sengi, at 32 cm (13 in) plus a 26 cm (10 in) tail, and generally eat insects, other invertebrates, and plants. The only macroscelid species with an International Union for Conservation of Nature (IUCN) population estimate is the golden-rumped sengi, listed as endangered with a population of around 13,000.

The nineteen extant species of Macroscelidea are grouped into a single family, Macroscelididae, and are split between six genera. Dozens of extinct macroscelid species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.

Conventions

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the IUCN Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the macroscelid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".

Classification

The order Macroscelidea consists of a single family, Macroscelididae, which contains twenty species divided into six genera. Many of these species are further subdivided into subspecies. This does not include hybrid species or extinct prehistoric species.

Family Macroscelididae

Genus Elephantulus (round-eared sengis): eight species
Genus Galegeeska (Horn of Africa sengis): two species
Genus Macroscelides (long-eared sengis): three species
Genus Petrodromus (four-toed sengi): one species
Genus Petrosaltator (North African sengi): one species
Genus Rhynchocyon (checkered sengis): four species

Macroscelididae

Rhynchocyon

Galegeeska

	Petrosaltator

	Petrodromus

Macroscelides

Elephantulus

Macroscelids

The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis.

Genus *Elephantulus* – Thomas, 1906 – eight species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Bushveld sengi	E. intufi (Smith, 1836)	Southern Africa	Size: 20–28 cm (8–11 in) long, plus 9–15 cm (4–6 in) tail Habitat: Savanna, shrubland, and grassland Diet: Ants, termites, and other small invertebrates, as well as plants	LC Unknown
Cape sengi	E. edwardii (Smith, 1839)	Southern South Africa	Size: 10–13 cm (4–5 in) long, plus 11–15 cm (4–6 in) tail Habitat: Shrubland and rocky areas Diet: Ants and termites, as well as other invertebrates	LC Unknown
Dusky sengi	E. fuscus (Peters, 1852)	Southern Africa	Size: 10–13 cm (4–5 in) long, plus 8–12 cm (3–5 in) tail Habitat: Savanna and shrubland Diet: Believed to be invertebrates	DD Unknown
Dusky-footed sengi	E. fuscipes (Thomas, 1894)	Central Africa	Size: 12–15 cm (5–6 in) long, plus 8–10 cm (3–4 in) tail Habitat: Savanna Diet: Believed to be invertebrates	DD Unknown
Eastern rock sengi	E. myurus Thomas, Schwann, 1906	Southern Africa	Size: 20–29 cm (8–11 in) long, plus 20–29 cm (8–11 in) tail Habitat: Savanna, grassland, and rocky areas Diet: Ants, termites, and other invertebrates, as well as plants	LC Unknown
Karoo rock sengi	E. pilicaudus Smit, Robinson, Watson, van Vuuren, 2008	Western South Africa	Size: 11–12 cm (4–5 in) long, plus 11–15 cm (4–6 in) tail Habitat: Shrubland and rocky areas Diet: Invertebrates	DD Unknown
Short-snouted sengi	E. brachyrhynchus (Smith, 1836)	Southern and southeastern Africa	Size: 11–13 cm (4–5 in) long, plus 8–11 cm (3–4 in) tail Habitat: Savanna and grassland Diet: Ants and termites, as well as small amounts of plants, fruit, and seeds	LC Unknown
Western rock sengi	E. rupestris (Smith, 1831)	Southern Africa	Size: 11–13 cm (4–5 in) long, plus 12–17 cm (5–7 in) tail Habitat: Savanna, shrubland, and rocky areas Diet: Invertebrates	LC Unknown

Genus *Galegeeska* – Heritage, Rayaleh, 2020 – two species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Somali sengi	G. revoili (Huet, 1881)	Horn of Africa	Size: 12–15 cm (5–6 in) long, plus 12–16 cm (5–6 in) tail Habitat: Savanna and shrubland Diet: Invertebrates as well as plants	DD Unknown
Rufous sengi	G. rufescens (Peters, 1878) Six subspecies G. r. boranus G. r. dundasi G. r. peasei G. r. pulcher G. r. rufescens G. r. somalicus	Eastern Africa	Size: 10–20 cm (4–8 in) long, plus 11–17 cm (4–7 in) tail Habitat: Savanna and grassland Diet: Termites and ants, as well as shoots, berries and roots	LC Unknown

Genus *Macroscelides* – Smith, 1829 – three species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Etendeka round-eared sengi	M. micus Dumbacher, Rathbun, 2014	Northwestern Namibia	Size: 8–10 cm (3–4 in) long, plus 8–10 cm (3–4 in) tail Habitat: Shrubland, grassland, and desert Diet: Invertebrates	LC Unknown
Namib round-eared sengi	M. flavicaudatus Lundholm, 1955	Namibia	Size: 10–12 cm (4–5 in) long, plus 9–14 cm (4–6 in) tail Habitat: Shrubland and desert Diet: Omnivorous, primarily invertebrates	LC Unknown
Round-eared sengi	M. proboscideus (Shaw, 1800)	Southern Africa	Size: 10–11 cm (4–4 in) long, plus 9–13 cm (4–5 in) tail Habitat: Shrubland and desert Diet: Termites, ants, and other small invertebrates, as well as plants	LC Unknown

Genus *Petrodromus* – Peters, 1846 – one species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Four-toed sengi	P. tetradactylus Peters, 1846 Nine subspecies P. t. beirae P. t. rovumae P. t. schwanni P. t. sultani P. t. swynnertoni P. t. tetradactylus P. t. tordayi P. t. warreni P. t. zanzibaricus	Central and southeastern Africa	Size: 19–23 cm (7–9 in) long, plus 15–17 cm (6–7 in) tail Habitat: Forest, savanna, and shrubland Diet: Termites, ants, and other small invertebrates, as well as plants	LC Unknown

Genus *Petrosaltator* – Rathbun, Dumbacher, 2016 – one species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
North African sengi	P. rozeti (Duvernoy, 1833) Two subspecies P. r. deserti P. r. rozeti	Northwestern Africa	Size: 11–13 cm (4–5 in) long, plus 13–16 cm (5–6 in) tail Habitat: Shrubland, rocky areas, and desert Diet: Believed to be ants, termites, and other invertebrates, as well as plants	LC Unknown

Genus *Rhynchocyon* – Peters, 1847 – four species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Black and rufous sengi	R. petersi Bocage, 1880 Two subspecies R. p. adersi R. p. petersi	Eastern Africa	Size: 25–31 cm (10–12 in) long, plus 25 cm (10 in) tail Habitat: Forest and shrubland Diet: Omnivorous; primarily ants, termites, and other invertebrates	LC Unknown
Checkered sengi	R. cirnei Peters, 1847 Six subspecies R. c. cirnei R. c. hendersoni R. c. macrurus R. c. reichardi R. c. shirensis R. c. stuhlmanni	Central and southeastern Africa	Size: 22–31 cm (9–12 in) long, plus 17–26 cm (7–10 in) tail Habitat: Forest and shrubland Diet: Invertebrates, as well as small mammals, amphibians, mollusks, birds, and bird eggs	LC Unknown
Golden-rumped sengi	R. chrysopygus Günther, 1881	Eastern Kenya	Size: 21–31 cm (8–12 in) long, plus 21–27 cm (8–11 in) tail Habitat: Forest and shrubland Diet: Wide variety of invertebrates	EN 13,000
Grey-faced sengi	R. udzungwensis Rovero, Rathbun, 2008	Central Tanzania	Size: 29–32 cm (11–13 in) long, plus 23–26 cm (9–10 in) tail Habitat: Forest Diet: Unknown	VU Unknown

References

↑ "Fossilworks: Macroscelidea". Paleobiology Database. University of Wisconsin–Madison. Retrieved February 27, 2022.
↑ Upham, N. S.; Esselstyn, J. A.; Jetz, W. (2019). "Inferring the mammal tree: Species-level sets of phylogenies for questions in ecology, evolution and conservation". PLOS Biology. 17 (12) e3000494. doi:10.1371/journal.pbio.3000494. PMC 6892540. PMID 31800571.
↑ Wilson, Reeder, pp. 82–85
1 2 Lindsey, David (2012). "Elephantulus intufi". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.
1 2 Rathbun, G. B. (2015). "Elephantulus intufi". IUCN Red List of Threatened Species. 2015 e.T42661A21289808. doi:10.2305/IUCN.UK.2015-2.RLTS.T42661A21289808.en.
1 2 Kingdon et al., p. 266
1 2 Rathbun, G. B.; Smit-Robinson, H. (2015). "Elephantulus edwardii". IUCN Red List of Threatened Species. 2015 e.T7136A21290344. doi:10.2305/IUCN.UK.2015-2.RLTS.T7136A21290344.en.
↑ Kingdon et al., p. 268
1 2 Rathbun, G. B. (2015). "Elephantulus fuscus". IUCN Red List of Threatened Species. 2015 e.T42660A21288491. doi:10.2305/IUCN.UK.2015-2.RLTS.T42660A21288491.en.
1 2 Kingdon et al., p. 259
↑ Kingdon et al., p. 267
1 2 Rathbun, G. B. (2015). "Elephantulus fuscipes". IUCN Red List of Threatened Species. 2015 e.T42659A21288575. doi:10.2305/IUCN.UK.2015-2.RLTS.T42659A21288575.en.
1 2 Jones, Jeremy (2002). "Elephantulus myurus". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.
1 2 Rathbun, G. B. (2015). "Elephantulus myurus". IUCN Red List of Threatened Species. 2015 e.T42662A21289491. doi:10.2305/IUCN.UK.2015-2.RLTS.T42662A21289491.en.
1 2 Smit, H. A.; Robinson, T. J.; Watson, J.; Jansen van Vuuren, B. (2008). "A New Species of Elephant-Shrew (Afrotheria: Macroscelidea: Elephantulus) from South Africa". Journal of Mammalogy. 89 (5): 1257–1269. doi:10.1644/07-MAMM-A-254.1. hdl:10019.1/117701.
1 2 Smit-Robinson, H.; Rathbun, G. (2015). "Elephantulus pilicaudus". IUCN Red List of Threatened Species. 2015 e.T165924A21291059. doi:10.2305/IUCN.UK.2015-2.RLTS.T165924A21291059.en.
↑ Kingdon et al., p. 265
1 2 Rathbun, G. B. (2015). "Elephantulus brachyrhynchus". IUCN Red List of Threatened Species. 2015 e.T42658A21288656. doi:10.2305/IUCN.UK.2015-2.RLTS.T42658A21288656.en.
↑ Gill, Elizabeth (2000). "Elephantulus brachyrhynchus". Animal Diversity Web. University of Michigan. Retrieved February 26, 2022.
1 2 Kingdon et al., p. 276
1 2 Rathbun, G. B.; Smit-Robinson, H. (2015). "Elephantulus rupestris". IUCN Red List of Threatened Species. 2015 e.T7138A21290631. doi:10.2305/IUCN.UK.2015-2.RLTS.T7138A21290631.en.
1 2 Heritage, S.; Rayaleh, H.; Awaleh, D. G.; Rathbun, G. B. (2020). "New records of a lost species and a geographic range expansion for sengis in the Horn of Africa". PeerJ. 8 e9652. doi:10.7717/peerj.9652. PMC 7441985. PMID 32879790.
1 2 Rathbun, G. B. (2017) [errata version of 2015 assessment]. "Elephantulus revoilii". IUCN Red List of Threatened Species. 2015 e.T7137A117060302. doi:10.2305/IUCN.UK.2015-2.RLTS.T7137A21290721.en.
↑ Kingdon et al., p. 275
1 2 Rathbun, G. B. (2015). "Elephantulus rufescens". IUCN Red List of Threatened Species. 2015 e.T42664A21289073. doi:10.2305/IUCN.UK.2015-2.RLTS.T42664A21289073.en.
↑ Awaad, Rania (2002). "Elephantulus rufescens". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.
1 2 Dumbacher, J. P.; Rathbun, G. B.; Osborne, T. O.; Griffin, M.; Eiseb, S. J. (2014). "A new species of round-eared sengi (genus Macroscelides) from Namibia". Journal of Mammalogy. 95 (3): 443–454. doi:10.1644/13-MAMM-A-159.
1 2 Rathbun, G. B.; Dumbacher, J. (2015). "Macroscelides micus". IUCN Red List of Threatened Species. 2015 e.T45434566A45436004. doi:10.2305/IUCN.UK.2015-2.RLTS.T45434566A45436004.en.
↑ Rathbun, G. B.; Dumbacher, J. P. (2015). "Home range and use of diurnal shelters by the Etendeka round-eared sengi, a newly discovered Namibian endemic desert mammal". PeerJ. 3 e1302. doi:10.7717/peerj.1302. PMC 4636418. PMID 26557433.
1 2 Dumbacher, J. P.; Rathbun, G. B.; Smit, H. A. (2012). "Phylogeny and Taxonomy of the Round-Eared Sengis or Elephant-Shrews, Genus Macroscelides (Mammalia, Afrotheria, Macroscelidea)". PLOS One. 7 (3) e32410. Bibcode:2012PLoSO...732410D. doi:10.1371/journal.pone.0032410. PMC 3314003. PMID 22479325.
1 2 Rathbun, G. B.; Eiseb, S. (2015). "Macroscelides flavicaudatus". IUCN Red List of Threatened Species. 2015 e.T45369877A45435876. doi:10.2305/IUCN.UK.2015-2.RLTS.T45369877A45435876.en.
1 2 Dohring, Alyce (2002). "Macroscelides proboscideus". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.
1 2 Rathbun, G. B.; Smit-Robinson, H. (2015). "Macroscelides proboscideus". IUCN Red List of Threatened Species. 2015 e.T45369602A45435551. doi:10.2305/IUCN.UK.2015-2.RLTS.T45369602A45435551.en.
1 2 Smith, Mary Alice (2002). "Petrodromus tetradactylus". Animal Diversity Web. University of Michigan. Retrieved February 26, 2022.
1 2 Rathbun, G. B.; FitzGibbon, C. (2015). "Petrodromus tetradactylus". IUCN Red List of Threatened Species. 2015 e.T42679A21290893. doi:10.2305/IUCN.UK.2015-2.RLTS.T42679A21290893.en.
1 2 Tao, Ran (2006). "Elephantulus rozeti". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.
1 2 Rathbun, G. B. (2015). "Elephantulus rozeti". IUCN Red List of Threatened Species. 2015 e.T42663A21289287. doi:10.2305/IUCN.UK.2015-2.RLTS.T42663A21289287.en.
1 2 Jacques, Abby (2013). "Rhynchocyon petersi". Animal Diversity Web. University of Michigan. Retrieved February 26, 2022.
1 2 Hoffmann, M.; Burgess, N.; Rovero, F. (2016). "Rhynchocyon petersi". IUCN Red List of Threatened Species. 2016 e.T19708A21286959. doi:10.2305/IUCN.UK.2016-1.RLTS.T19708A21286959.en.
1 2 Gasior, Robert (2006). "Rhynchocyon cirnei". Animal Diversity Web. University of Michigan. Retrieved February 26, 2022.
1 2 Hoffmann, M. (2020) [amended version of 2017 assessment]. "Rhynchocyon cirnei". IUCN Red List of Threatened Species. 2020 e.T19709A166489513. doi:10.2305/IUCN.UK.2020-1.RLTS.T19709A166489513.en.
↑ Kingdon et al., p. 284
1 2 FitzGibbon, C.; Rathbun, G. B. (2015). "Rhynchocyon chrysopygus". IUCN Red List of Threatened Species. 2015 e.T19705A21287265. doi:10.2305/IUCN.UK.2015-2.RLTS.T19705A21287265.en.
↑ Jansa, Sharon (1999). "Rhynchocyon chrysopygus". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.
1 2 Rovero, F.; Rathbun, G. B.; Perkins, A.; Jones, T.; Ribble, D.O.; Leonard, C.; Mwakisoma, R. R. (2008). "A new species of giant sengi or elephant-shrew (genus Rhynchocyon) highlights the exceptional biodiversity of the Udzungwa Mountains of Tanzania". Journal of Zoology. 274 (2): 126–133. Bibcode:2008JZoo..274..126R. doi:10.1111/j.1469-7998.2007.00363.x.
1 2 Rovero, F.; Rathbun, G. B. (2015). "Rhynchocyon udzungwensis". IUCN Red List of Threatened Species. 2015 e.T136309A21287423. doi:10.2305/IUCN.UK.2015-2.RLTS.T136309A21287423.en.

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[PDBMacroscelidea-1] "Fossilworks: Macroscelidea". Paleobiology Database. University of Wisconsin–Madison. Retrieved February 27, 2022.

[Classification1-2] Upham, N. S.; Esselstyn, J. A.; Jetz, W. (2019). "Inferring the mammal tree: Species-level sets of phylogenies for questions in ecology, evolution and conservation". PLOS Biology. 17 (12) e3000494. doi:10.1371/journal.pbio.3000494. PMC 6892540. PMID 31800571.

[MSW-3] Wilson, Reeder, pp. 82–85

[ADWBushveldelephantshrew-4] 1 2 Lindsey, David (2012). "Elephantulus intufi". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.

[IUCNBushveldelephantshrew-5] 1 2 Rathbun, G. B. (2015). "Elephantulus intufi". IUCN Red List of Threatened Species. 2015 e.T42661A21289808. doi:10.2305/IUCN.UK.2015-2.RLTS.T42661A21289808.en.

[MamAfrica266-6] 1 2 Kingdon et al., p. 266

[IUCNCapeelephantshrew-7] 1 2 Rathbun, G. B.; Smit-Robinson, H. (2015). "Elephantulus edwardii". IUCN Red List of Threatened Species. 2015 e.T7136A21290344. doi:10.2305/IUCN.UK.2015-2.RLTS.T7136A21290344.en.

[MamAfrica268-8] Kingdon et al., p. 268

[IUCNDuskyelephantshrew-9] 1 2 Rathbun, G. B. (2015). "Elephantulus fuscus". IUCN Red List of Threatened Species. 2015 e.T42660A21288491. doi:10.2305/IUCN.UK.2015-2.RLTS.T42660A21288491.en.

[MamAfrica259-10] 1 2 Kingdon et al., p. 259

[MamAfrica267-11] Kingdon et al., p. 267

[IUCNDusky-footedelephantshrew-12] 1 2 Rathbun, G. B. (2015). "Elephantulus fuscipes". IUCN Red List of Threatened Species. 2015 e.T42659A21288575. doi:10.2305/IUCN.UK.2015-2.RLTS.T42659A21288575.en.

[ADWEasternrockelephantshrew-13] 1 2 Jones, Jeremy (2002). "Elephantulus myurus". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.

[IUCNEasternrockelephantshrew-14] 1 2 Rathbun, G. B. (2015). "Elephantulus myurus". IUCN Red List of Threatened Species. 2015 e.T42662A21289491. doi:10.2305/IUCN.UK.2015-2.RLTS.T42662A21289491.en.

[Karoorockelephantshrewstudy-15] 1 2 Smit, H. A.; Robinson, T. J.; Watson, J.; Jansen van Vuuren, B. (2008). "A New Species of Elephant-Shrew (Afrotheria: Macroscelidea: Elephantulus) from South Africa". Journal of Mammalogy. 89 (5): 1257–1269. doi:10.1644/07-MAMM-A-254.1. hdl:10019.1/117701.

[IUCNKaroorockelephantshrew-16] 1 2 Smit-Robinson, H.; Rathbun, G. (2015). "Elephantulus pilicaudus". IUCN Red List of Threatened Species. 2015 e.T165924A21291059. doi:10.2305/IUCN.UK.2015-2.RLTS.T165924A21291059.en.

[MamAfrica265-17] Kingdon et al., p. 265

[IUCNShort-snoutedelephantshrew-18] 1 2 Rathbun, G. B. (2015). "Elephantulus brachyrhynchus". IUCN Red List of Threatened Species. 2015 e.T42658A21288656. doi:10.2305/IUCN.UK.2015-2.RLTS.T42658A21288656.en.

[ADWShort-snoutedelephantshrew-19] Gill, Elizabeth (2000). "Elephantulus brachyrhynchus". Animal Diversity Web. University of Michigan. Retrieved February 26, 2022.

[MamAfrica276-20] 1 2 Kingdon et al., p. 276

[IUCNWesternrockelephantshrew-21] 1 2 Rathbun, G. B.; Smit-Robinson, H. (2015). "Elephantulus rupestris". IUCN Red List of Threatened Species. 2015 e.T7138A21290631. doi:10.2305/IUCN.UK.2015-2.RLTS.T7138A21290631.en.

[Somalielephantshrewstudy-22] 1 2 Heritage, S.; Rayaleh, H.; Awaleh, D. G.; Rathbun, G. B. (2020). "New records of a lost species and a geographic range expansion for sengis in the Horn of Africa". PeerJ. 8 e9652. doi:10.7717/peerj.9652. PMC 7441985. PMID 32879790.

[IUCNSomalielephantshrew-23] 1 2 Rathbun, G. B. (2017) [errata version of 2015 assessment]. "Elephantulus revoilii". IUCN Red List of Threatened Species. 2015 e.T7137A117060302. doi:10.2305/IUCN.UK.2015-2.RLTS.T7137A21290721.en.

[MamAfrica275-24] Kingdon et al., p. 275

[IUCNRufouselephantshrew-25] 1 2 Rathbun, G. B. (2015). "Elephantulus rufescens". IUCN Red List of Threatened Species. 2015 e.T42664A21289073. doi:10.2305/IUCN.UK.2015-2.RLTS.T42664A21289073.en.

[ADWRufouselephantshrew-26] Awaad, Rania (2002). "Elephantulus rufescens". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.

[Etendekaround-earedsengistudy-27] 1 2 Dumbacher, J. P.; Rathbun, G. B.; Osborne, T. O.; Griffin, M.; Eiseb, S. J. (2014). "A new species of round-eared sengi (genus Macroscelides) from Namibia". Journal of Mammalogy. 95 (3): 443–454. doi:10.1644/13-MAMM-A-159.

[IUCNEtendekaround-earedsengi-28] 1 2 Rathbun, G. B.; Dumbacher, J. (2015). "Macroscelides micus". IUCN Red List of Threatened Species. 2015 e.T45434566A45436004. doi:10.2305/IUCN.UK.2015-2.RLTS.T45434566A45436004.en.

[Etendekaround-earedsengistudy2-29] Rathbun, G. B.; Dumbacher, J. P. (2015). "Home range and use of diurnal shelters by the Etendeka round-eared sengi, a newly discovered Namibian endemic desert mammal". PeerJ. 3 e1302. doi:10.7717/peerj.1302. PMC 4636418. PMID 26557433.

[Namibround-earedsengistudy-30] 1 2 Dumbacher, J. P.; Rathbun, G. B.; Smit, H. A. (2012). "Phylogeny and Taxonomy of the Round-Eared Sengis or Elephant-Shrews, Genus Macroscelides (Mammalia, Afrotheria, Macroscelidea)". PLOS One. 7 (3) e32410. Bibcode:2012PLoSO...732410D. doi:10.1371/journal.pone.0032410. PMC 3314003. PMID 22479325.

[IUCNNamibround-earedsengi-31] 1 2 Rathbun, G. B.; Eiseb, S. (2015). "Macroscelides flavicaudatus". IUCN Red List of Threatened Species. 2015 e.T45369877A45435876. doi:10.2305/IUCN.UK.2015-2.RLTS.T45369877A45435876.en.

[ADWRound-earedelephantshrew-32] 1 2 Dohring, Alyce (2002). "Macroscelides proboscideus". Animal Diversity Web. University of Michigan. Retrieved February 27, 2022.

[IUCNRound-earedelephantshrew-33] 1 2 Rathbun, G. B.; Smit-Robinson, H. (2015). "Macroscelides proboscideus". IUCN Red List of Threatened Species. 2015 e.T45369602A45435551. doi:10.2305/IUCN.UK.2015-2.RLTS.T45369602A45435551.en.

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[IUCNFour-toedelephantshrew-35] 1 2 Rathbun, G. B.; FitzGibbon, C. (2015). "Petrodromus tetradactylus". IUCN Red List of Threatened Species. 2015 e.T42679A21290893. doi:10.2305/IUCN.UK.2015-2.RLTS.T42679A21290893.en.

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