List of rodents

five rodent images
Clockwise from top-left: capybara (Hydrochoerus hydrochaeris), South African springhare (Pedetes capensis), golden-mantled ground squirrel (Callospermophilus lateralis), house mouse (Mus musculus), North American beaver (Castor canadensis)
Map of the world, with most of the world outside of Antarctica and the furthest northern arctic regions shaded blue
Worldwide distribution of rodent species

Rodentia is an order of placental mammals. Members of this order are called rodents. The order comprises 2,362 extant species, which are grouped into 513 genera. The largest order of mammals, rodents comprise about 40% of all mammal species worldwide. They are native to all major land masses except for Antarctica, and can be found worldwide. They live in a variety of habitats, particularly forests, grasslands, and shrublands, but also savannas, wetlands, deserts, and rocky areas. Most rodents are small animals with robust bodies, short limbs, and long tails. They range in length from the African pygmy mouse, at 4 cm (2 in) plus a 2 cm (1 in) tail, to the capybara, at 134 cm (53 in) plus a 2 cm (1 in) tail.

Rodentia is divided into five suborders: Anomaluromorpha, Castorimorpha, Hystricomorpha, Myomorpha, and Sciuromorpha. The suborders are further subdivided into clades and families. Anomaluromorpha contains two families, the scaly-tailed squirrels and the springhares. Castorimorpha is composed of three families and includes beavers, gophers, kangaroo rats, and pocket mice. There are 18 families in Hystricomorpha, which includes mole-rats, porcupines, cavies, hutias, tuco-tucos, and Neotropical spiny rats. Myomorpha contains nine families and includes jerboas, hamsters, voles, and a wide variety of rats and mice. The final three families of the order are in Sciuromorpha, which includes squirrels, chipmunks, and dormice. The exact organization of the species is not fixed, with many recent proposals made based on molecular phylogenetic analysis. In addition to the extant rodents, 38 species have been driven extinct since 1500 CE.

Conventions

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Range maps are provided wherever possible; if a range map is not available, a description of the collective range of species in that genera is provided. Ranges are based on the International Union for Conservation of Nature (IUCN) Red List of Threatened Species unless otherwise noted. All extinct genera or species listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".

Classification

The order Rodentia consists of 2,362 extant species belonging to 513 genera. This does not include hybrid species or extinct prehistoric species. Modern molecular studies indicate that the 511 genera can be grouped into 35 families; these families are divided between 5 named suborders and are grouped in those suborders into named clades, and some of these families are subdivided into named subfamilies. An additional 38 species have been driven extinct since 1500 CE: 8 in the family Capromyidae, 6 in Echimyidae, 11 in Cricetidae, and 13 in Muridae.

Suborder Anomaluromorpha

  • Family Anomaluridae (scaly-tailed squirrels): 3 genera, 7 species
  • Family Pedetidae (springhares): 1 genus, 2 species

Suborder Castorimorpha

Suborder Hystricomorpha

Suborder Myomorpha

  • Superfamily Dipodoidea
  • Superfamily Muroidea
    • Family Calomyscidae (brush-tailed mice): 1 genus, 8 species
    • Family Cricetidae (New World rats and mice)
      • Subfamily Arvicolinae (voles, lemmings, and muskrats): 30 genera, 158 species
      • Subfamily Cricetinae (hamsters): 9 genera, 18 species
      • Subfamily Neotominae (deer mice, packrats, and grasshopper mice): 16 genera, 124 species (1 extinct)
      • Subfamily Sigmodontinae (New World rats and mice): 87 genera, 395 species (10 extinct)
      • Subfamily Tylomyinae (vesper rats and climbing rats): 4 genera, 10 species
    • Family Muridae (Old World rats and mice)
      • Subfamily Deomyinae (spiny mice and brush-furred rats): 4 genera, 55 species
      • Subfamily Gerbillinae (gerbils, jirds, and sand rats): 14 genera, 102 species
      • Subfamily Leimacomyinae (Togo mouse): 1 genus, 1 species
      • Subfamily Lophiomyinae (maned rat): 1 genus, 1 species
      • Subfamily Murinae (Old World mice and rats): 136 genera, 655 species (13 extinct)
    • Family Nesomyidae (African and Malagasy rats and mice)
      • Subfamily Cricetomyinae (hamster-rats and pouched rats): 3 genera, 8 species
      • Subfamily Delanymyinae (Delany's mouse): 1 genus, 1 species
      • Subfamily Dendromurinae (climbing mice and fat mice): 6 genera, 26 species
      • Subfamily Mystromyinae (white-tailed rat): 1 genus, 1 species
      • Subfamily Nesomyinae (Malagasy rodents): 9 genera, 27 species
      • Subfamily Petromyscinae (rock mice): 1 genus, 4 species
    • Family Platacanthomyidae (Oriental dormice): 2 genera, 2 species
    • Family Spalacidae (mole-rats)
      • Subfamily Myospalacinae (zokors): 2 genera, 6 species
      • Subfamily Rhizomyinae (bamboo rats and mole-rats): 3 genera, 6 species
      • Subfamily Spalacinae (blind mole-rats): 2 genera, 11 species

Suborder Sciuromorpha

  • Family Aplodontiidae (mountain beaver): 1 genus, 1 species
  • Family Gliridae (dormice)
    • Subfamily Glirinae (Japanese dormice and edible dormice): 2 genera, 2 species
    • Subfamily Graphiurinae (African dormice): 1 genus, 15 species
    • Subfamily Leithiinae (dormice): 6 genera, 12 species
  • Family Sciuridae (squirrels, chipmunks, and marmots)
    • Subfamily Callosciurinae (Asian squirrels): 14 genera, 67 species
    • Subfamily Ratufinae (giant squirrels): 1 genus, 4 species
    • Subfamily Sciurillinae (Neotropical pygmy squirrel): 1 genus, 1 species
    • Subfamily Sciurinae (tree squirrels and flying squirrels): 20 genera, 2 species
    • Subfamily Xerinae (ground squirrels, chipmunks, and prairie dogs): 24 genera, 2 species
Rodentia

Rodents

The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.

Suborder Anomaluromorpha

Family Anomaluridae

Members of the Anomaluridae family are called anomalurids or scaly-tailed squirrels. They eat a variety of vegetation, and some also eat insects. Anomaluridae comprises seven extant species, divided into three genera.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Anomalurus
(scaly-tailed flying squirrel)

Brown squirrel

Gervais, 1849

Four species
Sub-Saharan Africa (in red)
Map of range
Size range: 19 cm (7 in) long, plus 12 cm (5 in) tail (dwarf scaly-tailed squirrel) to 46 cm (18 in) long, plus 45 cm (18 in) tail (Pel's flying squirrel)

Habitats: Forest

Diets: Bark and fruit as well as flowers, leaves, nuts, and insects
Idiurus
(flying mice)

Brown squirrel

Matschie, 1894

Two species
Central and western Africa (in blue)
Map of range
Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (pygmy scaly-tailed flying squirrel) to 12 cm (5 in) long, plus 19 cm (7 in) tail (long-eared flying mouse)

Habitats: Forest

Diets: Fruit
Zenkerella

Gray squirrel

Matschie, 1898

One species
West-central Africa (in purple)
Map of range
Size range: 18–23 cm (7–9 in) long, plus 15–18 cm (6–7 in) tail

Habitats: Forest and savanna

Diets: Vegetation

Family Pedetidae

Members of the Pedetidae family are called pedetids or springhares. They eat bulbs and roots, as well as stems and grain. Pedetidae comprises two extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Pedetes
(springhare)

Brown springhare

Illiger, 1811

Two species
Southern and eastern Africa
Map of range
Size range: 34–46 cm (13–18 in) long, plus 39–48 cm (15–19 in) tail

Habitats: Savanna, shrubland, and grassland

Diets: Bulbs and roots, as well as stems and grain

Suborder Castorimorpha

Superfamily Castoroidea

Family Castoridae

Members of the Castoridae family are called castorids or beavers. They eat woody and aquatic plants. Castoridae comprises two extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Castor
(beaver)

Brown springhare

Linnaeus, 1758

Two species
North America, Europe, and Asia
Map of range
Size range: 80–90 cm (31–35 in) long, plus 20–30 cm (8–12 in) tail

Habitats: Forest, shrubland, and inland wetlands

Diets: Bark, twigs, leaves, aquatic plants, and tree and shrub roots

Superfamily Geomyoidea

Family Geomyidae

Members of the Geomyidae family are called geomyids, pocket gophers, or gophers. They primarily eat roots, bulbs, and rhizomes, as well as other vegetation. Geomyidae comprises 36 extant species in 7 genera.

Not assigned to a named subfamily – seven genera
Name Authority and species Range Size and ecology
Cratogeomys
(pocket gopher)

Brown gopher

Merriam, 1895

Seven species
Southern United States and Mexico
Map of range
Size range: 14 cm (6 in) long, plus 5 cm (2 in) tail (Goldman's pocket gopher) to 28 cm (11 in) long, plus 13 cm (5 in) tail (Merriam's pocket gopher)

Habitats: Grassland, shrubland, forest, and desert

Diets: A variety of roots and vegetation, including cacti
Geomys
(eastern pocket gopher)

Brown gopher

Rafinesque, 1817

Nine species
South-central Canada, United States, and Mexico
Map of range
Size range: 11 cm (4 in) long, plus 5 cm (2 in) tail (Knox Jones's pocket gopher) to 28 cm (11 in) long, plus 12 cm (5 in) tail (Texas pocket gopher)

Habitats: Grassland, shrubland, savanna, forest, and desert

Diets: Roots, rhizomes, and bulbs, as well as other vegetation
Heterogeomys
(pocket gopher)
Le Conte, 1852

Three species
Central America and Central Mexico
Map of range
Size range: 18 cm (7 in) long, plus 8 cm (3 in) tail (Cherrie's pocket gopher) to 29 cm (11 in) long, plus 10 cm (4 in) tail (big pocket gopher)

Habitats: Forest

Diets: A variety of vegetation
Orthogeomys
(taltuza)

Brown gopher head

Merriam, 1895

Six species
Mexico and Central America
Map of range
Size range: 18 cm (7 in) long, plus 8 cm (3 in) tail (Underwood's pocket gopher) to 30 cm (12 in) long, plus 14 cm (6 in) tail (giant pocket gopher)

Habitats: Intertidal marine, forest, and grassland

Diets: A variety of vegetation
Pappogeomys Merriam, 1895

One species
Western-central Mexico
Map of range
Size: 12–18 cm (5–7 in) long, plus 6–10 cm (2–4 in) tail

Habitats: Forest, shrubland, and grassland

Diet: Roots and other vegetation
Thomomys
(smooth-toothed pocket gopher)

Brown gopher

Wied-Neuwied, 1839

Nine species
Southwestern Canada, western United States, and northern Mexico
Map of range
Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (Botta's pocket gopher) to 24 cm (9 in) long, plus 10 cm (4 in) tail (Camas pocket gopher)

Habitats: Inland wetlands, grassland, shrubland, savanna, forest, and desert

Diets: Roots, bulbs, leaves, and other vegetation
Zygogeomys Merriam, 1895

One species
Central Mexico
Map of range
Size: 17–24 cm (7–9 in) long, plus 6–10 cm (2–4 in) tail

Habitats: Forest

Diet: Roots and rhizomes
Family Heteromyidae

Members of the Heteromyidae family are called heteromyids and include kangaroo rats, kangaroo mice, pocket mice, and spiny pocket mice. They primarily eat seeds, leaves, and other vegetation, as well as grain and insects. Heteromyidae comprises 63 extant species in 5 genera. These genera are divided between three subfamilies: Dipodomyinae, containing the kangaroo rats and kangaroo mice; Heteromyinae, or the spiny pocket mice; and Perognathinae, or the pocket mice.

Subfamily Dipodomyinae – Gervais, 1853 – two genera
Name Authority and species Range Size and ecology
Dipodomys
(kangaroo rat)

Brown and white kangaroo rat

Gray, 1841

20 species
Southern Canada, western United States, and Mexico
Map of range
Size range: 7 cm (3 in) long, plus 13 cm (5 in) tail (Fresno kangaroo rat) to 16 cm (6 in) long, plus 21 cm (8 in) tail (desert kangaroo rat)

Habitats: Rocky areas, shrubland, savanna, grassland, coastal marine, desert, and forest

Diets: Seeds, as well as fruit, leaves, stems, buds, and insects
Microdipodops
(kangaroo mouse)

Brown and white long-tailed rat

Merriam, 1891

Two species
Western United States
Map of range
Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail to 7 cm (3 in) long, plus 11 cm (4 in) tail (dark kangaroo mouse)

Habitats: Desert and shrubland

Diets: Grains and insects
Subfamily Heteromyinae – Gray, 1868 – one genus
Name Authority and species Range Size and ecology
Heteromys
(spiny pocket mouse)

Gray spiny pocket mouse

Desmarest, 1817

Southern United States, Mexico, Central America, and northeastern South America
Map of range
Size range: 10 cm (4 in) long, plus 12 cm (5 in) tail (Jaliscan spiny pocket mouse) to 17 cm (7 in) long, plus 20 cm (8 in) tail (Nelson's spiny pocket mouse)

Habitats: Shrubland and forest

Diets: Seeds, leaves, twigs, and succulent plants
Subfamily Perognathinae – Coues, 1875 – two genera
Name Authority and species Range Size and ecology
Chaetodipus
(coarse-haired pocket mouse)

Gray pocket mouse

Merriam, 1889

Eighteen species
United States and Mexico
Map of range
Size range: 6 cm (2 in) long, plus 8 cm (3 in) tail (little desert pocket mouse) to 12 cm (5 in) long, plus 12 cm (5 in) tail (Hispid pocket mouse)

Habitats: Rocky areas, shrubland, grassland, desert, and forest

Diets: Seeds, as well as vegetation and insects
Perognathus
(silky pocket mouse)

Brown pocket mouse

Wied-Neuwied, 1839

Nine species
Southern Canada, United States, and Mexico
Map of range
Size range: 5 cm (2 in) long, plus 4 cm (2 in) tail (Merriam's pocket mouse) to 8 cm (3 in) long, plus 9 cm (4 in) tail (white-eared pocket mouse)

Habitats: Desert, shrubland, savanna, grassland, and forest

Diets: Seeds, as well as vegetation and insects

Suborder Hystricomorpha

Parvorder Caviomorpha

Superfamily Cavioidea
Family Caviidae

Members of the Caviidae family are called caviids and include guinea pigs, cavies, maras, and capybaras. They are herbivores and eat a wide variety of vegetation, primarily grass and leaves. Caviidae comprises 21 extant species which are divided into 6 genera in 3 subfamilies: Caviinae, containing the guinea pigs and cavies; Dolichotinae, containing the maras; and Hydrochoerinae, containing rock cavies and capybaras.

Subfamily Caviinae – Fischer de Waldheim, 1817 – three genera
Name Authority and species Range Size and ecology
Cavia
(guinea pig)

Black and white guinea pig

Pallas, 1766

Seven species
South America (guinea pig domesticated worldwide)
Map of range
Size range: 20 cm (8 in) long, with no tail (guinea pig) to 39 cm (15 in) long, with no tail (Brazilian guinea pig)

Habitats: Grassland, forest, rocky areas, savanna, and inland wetlands

Diets: Wide variety of vegetation
Galea
(yellow-toothed cavy)

Brown cavy

Meyen, 1832

Five species
South America
Map of range
Size range: 19 cm (7 in) long, with no tail (lowland yellow-toothed cavy) to 24 cm (9 in) long, with no tail (southern highland yellow-toothed cavy)

Habitats: Savanna, rocky areas, shrubland, and grassland

Diets: Grass, forbs, and other vegetation
Microcavia
(mountain cavy)

Brown cavy

Gervais & Ameghino, 1880

Three species
Southern and western South America
Map of range
Size range: 19–20 cm (7–8 in) long, with no tail (Andean mountain cavy)

Habitats: Desert, grassland, forest, rocky areas, savanna, and inland wetlands

Diets: Leaves, as well as fruit
Subfamily Dolichotinae – Pocock, 1922 – one genus
Name Authority and species Range Size and ecology
Dolichotis
(mara)

Brown long-legged cavy

Desmarest, 1820

Two species
Southern and western South America
Map of range
Size range: 42 cm (17 in) long, plus 1 cm (0.4 in) tail (Chacoan mara) to 80 cm (31 in) long, plus 4 cm (2 in) tail (Patagonian mara)

Habitats: Savanna, forest, shrubland, and grassland

Diets: Wide variety of vegetation
Subfamily Hydrochoerinae – J. E. Gray, 1825 – two genera
Name Authority and species Range Size and ecology
Hydrochoerus
(capybara)

Brown capybara

Brisson, 1762

Two species
Northwestern South America and Panama and South America
Map of range
Size range: 102 cm (40 in) long, plus 1 cm (0.4 in) tail (lesser capybara) to 134 cm (53 in) long, plus 2 cm (0.8 in) tail (capybara)

Habitats: Forest, inland wetlands, and savanna

Diets: Grass, as well as aquatic plants, grains, melons, and squashes
Kerodon
(rock cavy)

Two brown cavies

F. Cuvier, 1825

Two species
Eastern Brazil
Map of range
Size range: 30 cm (12 in) long, with no tail (rock cavy) to 38 cm (15 in) long, with no tail (acrobatic cavy)

Habitats: Forest, rocky areas, and savanna

Diets: Leaves, flowers, buds, bark, and occasionally fruit
Family Cuniculidae

Members of the Cuniculidae family are called cuniculids or pacas. It contains two species, which eat leaves, stems, roots, seeds, and fruit.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Cuniculus
(paca)

Brown paca

Brisson, 1762

Two species
Mexico, Cuba, Central America, and South America
Map of range
Size range: 50 cm (20 in) long, plus 1 cm (0.4 in) tail (lowland paca) to 80 cm (31 in) long, plus 3 cm (1 in) tail (mountain paca)

Habitats: Forest and inland wetlands

Diets: Leaves, stems, roots, seeds, and fruit
Family Dasyproctidae

Members of the Dasyproctidae family are called dasyproctids and include agoutis and acouchis. They are herbivores and primarily eat fruit, vegetables, and succulent plants. Dasyproctidae comprises fifteen extant species which are divided into two genera.

Not assigned to a named subfamily – two genera
Name Authority and species Range Size and ecology
Dasyprocta
(agouti)

Brown agouti

Bonaparte, 1838

Thirteen species
Mexico, Cuba, Central America, and South America (in red)
Map of range
Size range: 43 cm (17 in) long, plus 1 cm (0.4 in) tail (Azara's agouti) to 76 cm (30 in) long, plus 4 cm (2 in) tail (black agouti)

Habitats: Savanna and forest

Diets: Fruit, seeds, vegetables, and succulent plants, as well as crabs
Myoprocta
(achouchi)

Brown achouchi

Thomas, 1903

Two species
Northern South America (in blue)
Map of range
Size range: 30 cm (12 in) long, plus 4 cm (2 in) tail (green acouchi) to 39 cm (15 in) long, plus 8 cm (3 in) tail (red acouchi)

Habitats: Forest

Diets: Fruit, seeds, vegetables, and succulent plants
Superfamily Chinchilloidea
Family Chinchillidae

Members of the Chinchillidae family are called chinchillids and contain chinchillas and viscachas. They are herbivores and eat grass and seeds, as well as other vegetation. Chinchillidae comprises six extant species which are divided into three genera, as well as a single extinct species.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Chinchilla
(chinchilla)

Gray chinchilla

Bennett, 1829

Two species
Western South America
Map of range
Size range: 22 cm (9 in) long, plus 14 cm (6 in) tail (long-tailed chinchilla) to 38 cm (15 in) long, plus 10 cm (4 in) tail (short-tailed chinchilla)

Habitats: Grassland, shrubland, and rocky areas

Diets: A wide variety of vegetation
Lagidium
(viscacha)

Brown viscacha

Meyen, 1833

Three species
Western and southern South America
Map of range
Size range: 29 cm (11 in) long, plus 31 cm (12 in) tail (southern viscacha) to 48 cm (19 in) long, plus 31 cm (12 in) tail (Wolffsohn's viscacha)

Habitats: Rocky areas

Diets: A wide variety of vegetation, as well as lichen and moss
Lagostomus
(plains viscacha)

Gray viscacha

Brookes, 1828

Two species
Southern South America
Map of range
Size range: 39–61 cm (15–24 in) long, plus 13–20 cm (5–8 in) tail (plains viscacha)

Habitats: Savanna, grassland, and desert

Diets: Grass and seeds, as well as other vegetation
Family Dinomyidae

Members of the Dinomyidae family are called dinomyids. It contains a single species, which eats fruit, leaves, and stems.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Dinomys

Brown pacarana

Peters, 1873

One species
Northwestern South America
Map of range
Size: 73–79 cm (29–31 in) long, plus 14–23 cm (6–9 in) tail (pacarana)

Habitats: Forest and grassland

Diet: Fruit, leaves, and stems
Superfamily Erethizontoidea
Family Erethizontidae

Members of the Erethizontidae family are called erethizontids or New World porcupines. They are herbivores and primarily eat leaves, stems, fruit, flowers, and roots. Erethizontidae comprises eighteen extant species in three genera, grouped into two subfamilies: Chaetomyinae, containing a single species, and Erethizontinae, containing the other seventeen.

Subfamily Chaetomyinae – Thomas, 1897 – one genus
Name Authority and species Range Size and ecology
Chaetomys

Brown porcupine

J. E Gray, 1843

One species
Eastern Brazil
Map of range
Size: 36–45 cm (14–18 in) long, plus 26–27 cm (10–11 in) tail

Habitats: Forest

Diet: Nuts
Subfamily Erethizontinae – Bonaparte, 1845 – two genera
Name Authority and species Range Size and ecology
Coendou
(prehensile-tailed porcupine)

Black and yellow porcupine

Lacépède, 1799

Sixteen species
Mexico, Central America, and South America
Map of range
Size range: 24 cm (9 in) long, plus 20 cm (8 in) tail (Paraguaian hairy dwarf porcupine) to 55 cm (22 in) long, plus 47 cm (19 in) tail (Baturite porcupine)

Habitats: Forest and savanna

Diets: Leaves, stems, fruit, flowers, and roots
Erethizon

Brown and white porcupines

F. Cuvier, 1823

One species
North America
Map of range
Size: 60–130 cm (24–51 in) long, plus 16–25 cm (6–10 in) tail

Habitats: Forest, shrubland, and grassland

Diet: Buds, twigs, roots, stems, leaves, flowers, seeds, berries, nuts, and other vegetation
Superfamily Octodontoidea
Family Abrocomidae

Members of the Abrocomidae family are called abrocomids or chinchilla rats. They are herbivores and eat a variety of vegetation. Abrocomidae comprises ten extant species which are divided into two genera.

Not assigned to a named subfamily – two genera
Name Authority and species Range Size and ecology
Abrocoma
(chinchilla rat)

Brown chinchilla rat

Waterhouse, 1837

Eight species
Western South America (in red)
Map of range
Size range: 15 cm (6 in) long, plus 5 cm (2 in) tail (ashy chinchilla rat) to 23 cm (9 in) long, plus 28 cm (11 in) tail (Bennett's chinchilla rat)

Habitats: Shrubland, inland wetlands, forest, rocky areas, grassland, and savanna

Diets: A variety of vegetation
Cuscomys
(arboreal chinchilla rat)

Drawing of chinchilla rat

Emmons, 1999

Two species
Peru
Map of range
Size range: About 35 cm (14 in) long, plus about 26 cm (10 in) tail (Asháninka arboreal chinchilla rat)

Habitats: Forest

Diets: A variety of vegetation
Family Capromyidae

Members of the Capromyidae family are called capromyids or hutias. They are omnivores and eat leaves, fruit, bark, and small animals. Capromyidae comprises ten extant species in five genera, grouped into two subfamilies: Capromyinae, containing nine species, and Plagiodontinae, containing a single species. Additionally, two species in Capromyinae, three species in Plagiodontinae, the sole species in the subfamily Hexolobodontinae, and the two species in the subfamily Isolobodontinae were driven extinct after 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s.

Subfamily Capromyinae – Smith, 1842 – four genera
Name Authority and species Range Size and ecology
Capromys

Brown hutia

Desmarest, 1822

One species
Cuba
Map of range
Size: 30–62 cm (12–24 in) long, plus 13–31 cm (5–12 in) tail

Habitats: Forest and caves

Diet: Leaves, fruit, bark, lizards, and other small animals
Geocapromys
(Bahamian and Jamaican hutia)

Stuffed brown hutia

Chapman, 1901

Four species
Caribbean
Map of range
Size range: 28 cm (11 in) long, plus 6 cm (2 in) tail (Bahamian hutia) to 45 cm (18 in) long, plus 7 cm (3 in) tail (Jamaican coney)

Habitats: Shrubland, intertidal marine, forest, and rocky areas

Diets: Bark, twigs, and leaves
Mesocapromys
(small Cuban hutia)

Brown hutia

Varona, 1970

Five species
Cuba
Map of range
Size range: 20 cm (8 in) long, plus 17 cm (7 in) tail (Dwarf hutia) to 38 cm (15 in) long, plus 32 cm (13 in) tail (Black-tailed hutia)

Habitats: Inland wetlands and forest

Diets: Vegetation and small animals
Mysateles

Stuffed brown hutia

Lesson, 1842

One species
Western Cuba
Map of range
Size: 33–45 cm (13–18 in) long, plus 25–35 cm (10–14 in) tail

Habitats: Forest

Diet: Leaves
Subfamily Hexolobodontinae † – Woods, 1989 – one genus
Name Authority and species Range Size and ecology
Hexolobodon †

Hutia jawbone

Miller, 1929

One species
Hispaniola Size: Unknown

Habitats: Unknown

Diet: Unknown
Subfamily Isolobodontinae † – Woods, 1989 – one genus
Name Authority and species Range Size and ecology
Isolobodon †
(laminar-toothed hutia)

Hutia jawbone

J. A. Allen, 1916

Two species
Hispaniola, Puerto Rico, and nearby islands Size range: Unknown

Habitats: Forest

Diets: Unknown
Subfamily Plagiodontinae – Ellerman, 1940 – three genera
Name Authority and species Range Size and ecology
Hyperplagiodontia † Rímoli, 1977

One species
Hispaniola Size: Unknown

Habitats: Unknown

Diet: Bark, leaves, and tree buds
Plagiodontia
(Hispaniolan hutia)

Drawing of black hutia

F. Cuvier, 1836

Two species
Hispaniola
Map of range
Size range: 31–40 cm (12–16 in) long, plus 12–16 cm (5–6 in) tail (Hispaniolan hutia)

Habitats: Rocky areas, and forest

Diets: Bark, leaves, and tree buds
Rhizoplagiodontia † Woods, 1989

One species
Hispaniola Size: Unknown

Habitats: Unknown

Diet: Unknown
Family Ctenomyidae

Members of the Ctenomyidae family are called ctenomyids or tuco-tucos. They are herbivores and eat a wide variety of vegetation. Ctenomyidae comprises 61 extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Ctenomys
(tuco-tuco)

Brown rodent

Blainville, 1826

61 species
Southern South America
Map of range
Size range: 11 cm (4 in) long, plus 7 cm (3 in) tail (white-toothed tuco-tuco) to 33 cm (13 in) long, plus 14 cm (6 in) tail (Conover's tuco-tuco)

Habitats: Forest, desert, grassland, inland wetlands, coastal marine, shrubland, and savanna

Diets: A wide variety of vegetation
Family Echimyidae

Members of the Echimyidae family are called echimyids or Neotropical spiny rats, and include spiny rats, tree-rats, and cave rats. They primarily eat a wide variety of vegetation, though some species also eat insects. Echimyidae comprises 88 extant species in 23 genera, split into three subfamilies: Caterodontinae, containing a single species; Echimyinae, containing 75 species, and Euryzygomatomyinae, containing 12 species. Additionally, six species of Caribbean cave rat, comprising the three genera of the subfamily Heteropsomyinae, were driven to extinction since 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s.

Subfamily Carterodontinae – Courcelle, Tilak, Leite, Douzery, & Fabre, 2019 – one genus
Name Authority and species Range Size and ecology
Carterodon Waterhouse, 1848

One species
Eastern Brazil
Map of range
Size: 13–25 cm (5–10 in) long, plus 6–10 cm (2–4 in) tail

Habitats: Forest and savanna

Diet: Vegetation
Subfamily Echimyinae – Gray, 1825 – eighteen genera
Name Authority and species Range Size and ecology
Callistomys

Gray and white spiny rat

Emmons & Vucetich, 1998

One species
Eastern Brazil
Map of range
Size: 25–29 cm (10–11 in) long, plus 27–32 cm (11–13 in) tail

Habitats: Forest

Diet: Vegetation
Dactylomys
(bamboo rat)

Drawing of gray spiny rat

Geoffroy, 1838

Three species
Northern and western South America
Map of range
Size range: 24 cm (9 in) long, plus 32 cm (13 in) tail (montane bamboo rat) to 31 cm (12 in) long, plus 39 cm (15 in) tail (Amazon bamboo rat)

Habitats: Forest

Diets: Bamboo and other plants
Diplomys
(soft-furred spiny rat)

Drawing of brown spiny rat

Thomas, 1916

Two species
Northwestern South America and Panama
Map of range
Size range: 21–39 cm (8–15 in) long, plus 17–27 cm (7–11 in) tail (Colombian soft-furred spiny rat)

Habitats: Forest

Diets: Seeds and fruit
Echimys
(spiny tree-rat)

Brown and white spiny rat

F. Cuvier, 1809

Three species
Northern South America
Map of range
Size range: 24 cm (9 in) long, plus 34 cm (13 in) tail (Vieira's tree-rat) to 33 cm (13 in) long, plus 38 cm (15 in) tail (dark spiny tree-rat)

Habitats: Forest

Diets: Vegetation
Hoplomys

Brown spiny rat

J. A. Allen, 1908

One species
Central America and northwestern South America
Map of range
Size: 21–32 cm (8–13 in) long, plus 11–24 cm (4–9 in) tail

Habitats: Forest

Diet: Wide variety of vegetation, as well as insects
Isothrix
(brush-tailed rat)

Brown spiny rat

Wagner, 1845

Six species
Northern and western South America
Map of range
Size range: 17 cm (7 in) long, plus 17 cm (7 in) tail (Plain brush-tailed rat) to 29 cm (11 in) long, plus 27 cm (11 in) tail (Rio Negro brush-tailed rat)

Habitats: Forest and grassland

Diets: Vegetation
Kannabateomys

Brown and white spiny rat

Jentink, 1891

One species
Eastern South America
Map of range
Size: 23–35 cm (9–14 in) long, plus 30–42 cm (12–17 in) tail

Habitats: Forest

Diet: Bamboo
Lonchothrix Thomas, 1920

One species
Northern Brazil
Map of range
Size: 15–22 cm (6–9 in) long, plus 15–23 cm (6–9 in) tail

Habitats: Forest and grassland

Diet: Vegetation
Makalata
(armored tree-rat)

Brown spiny rat

Husson, 1978

Three species
Northern South America
Map of range
Size range: 15 cm (6 in) long, plus 15 cm (6 in) tail (Brazilian spiny tree-rat) to 28 cm (11 in) long, plus 24 cm (9 in) tail (long-tailed armored tree-rat)

Habitats: Forest

Diets: Fruit and seeds
Mesomys
(spiny tree-rat)

Brown spiny rat

Wagner, 1845

Four species
Northern South America
Map of range
Size range: 15–20 cm (6–8 in) long, plus 12–20 cm (5–8 in) tail (Pará spiny tree-rat)

Habitats: Forest

Diets: Vegetation
Myocastor

Brown nutria in water

Kerr, 1792

One species
Southern South America, and introduced in United States, Europe, and Japan
Map of range
Size: 47–57 cm (19–22 in) long, plus 34–40 cm (13–16 in) tail

Habitats: Inland wetlands

Diet: Roots and aquatic plants
Olallamys
(olalla rat)

Drawing of brown and white spiny rat

Emmons, 1988

Two species
Colombia and western Venezuela
Map of range
Size range: 15 cm (6 in) long, plus 25 cm (10 in) tail (white-tailed olalla rat) to 22 cm (9 in) long, plus 34 cm (13 in) tail (greedy olalla rat)

Habitats: Forest and shrubland

Diets: Vegetation
Pattonomys
(speckled tree-rat)
Emmons, 2005

Five species
Colombia, Venezuela, and Ecuador
Map of range
Size range: 20 cm (8 in) long, plus 21 cm (8 in) tail (speckled spiny tree-rat) to 27 cm (11 in) long, plus 23 cm (9 in) tail (Orinocoan speckled tree-rat)

Habitats: Forest and savanna

Diets: Vegetation
Phyllomys
(Atlantic tree-rat)

Brown spiny rat

Lund, 1839

Southern and eastern Brazil
Map of range
Size range: 17 cm (7 in) long, plus 19 cm (7 in) tail (golden Atlantic tree-rat) to 30 cm (12 in) long, plus 34 cm (13 in) tail (giant Atlantic tree-rat)

Habitats: Forest

Diets: Vegetation
Proechimys
(spiny rat)

Brown spiny rat

Allen, 1899

22 species
Central America and Northern and western South America
Map of range
Size range: 14 cm (6 in) long, plus 9 cm (4 in) tail (Kulina spiny rat) to 30 cm (12 in) long, plus 20 cm (8 in) tail (Pacific spiny rat)

Habitats: Forest and savanna

Diets: Wide variety of vegetation
Santamartamys

Red, brown, and white spiny rat

Emmons, 2005

One species
Northern Colombia
Map of range
Size: About 19 cm (7 in)

Habitats: Unknown

Diet: Seeds and fruit
Thrichomys
(punaré)

Gray spiny rat

Trouessart, 1880

Four species
Southern and eastern Brazil
Map of range
Size range: 12–24 cm (5–9 in) long, plus 13–21 cm (5–8 in) tail (Sao Lourenço punaré)

Habitats: Forest, rocky areas, savanna, and shrubland

Diets: Seeds, fruit, cacti, and insects
Toromys
(giant tree-rat)
Iack-Ximenes, Vivo, & Percequillo, 2005

Two species
Northern Brazil and Peru
Map of range
Size range: 21 cm (8 in) long, plus 18 cm (7 in) tail (Peruvian toro) to 30 cm (12 in) long, plus 28 cm (11 in) tail (giant tree-rat)

Habitats: Forest

Diets: Vegetation
Subfamily Euryzygomatomyinae – Fabre, Upham, Emmons, Justy, Leite, Loss, Orlando, Tilak, Patterson, & Douzery, 2017 – three genera
Name Authority and species Range Size and ecology
Clyomys Thomas, 1916

One species
Paraguay and southern Brazil
Map of range
Size: 15–29 cm (6–11 in) long, plus 4–9 cm (2–4 in) tail

Habitats: Savanna

Diet: Vegetation
Euryzygomatomys

Drawing of brown spiny rat

Goeldi, 1901

One species
Southern Brazil and northeastern Argentina
Map of range
Size: 16–20 cm (6–8 in) long, plus 6–7 cm (2–3 in) tail

Habitats: Forest and grassland

Diet: Vegetation
Trinomys
(Atlantic spiny rat)

Brown and white spiny rat

Thomas, 1921

Eastern Brazil
Map of range
Size range: 14 cm (6 in) long, plus 16 cm (6 in) tail (Yonenaga's Atlantic spiny rat) to 27 cm (11 in) long, plus 23 cm (9 in) tail (spiked Atlantic spiny rat)

Habitats: Forest, savanna, rocky areas, grassland, and shrubland

Diets: Wide variety of vegetation
Subfamily Heteropsomyinae † – Anthony, 1917 – three genera
Name Authority and species Range Size and ecology
Boromys †
(Cuban cave rat)
Miller, 1916

Two species
Cuba Size range: Unknown

Habitats: Unknown

Diets: Unknown
Brotomys †
(edible rat)
Miller, 1916

Two species
Dominican Republic and Haiti Size range: Unknown

Habitats: Unknown

Diets: Unknown
Heteropsomys †
(Puerto Rican cave rat)
Anthony, 1916

Two species
Puerto Rico Size range: Unknown

Habitats: Unknown

Diets: Unknown
Family Octodontidae

Members of the Octodontidae family are called octodontids and include degus, rock rats, and viscacha rats. They are herbivores, and eat a variety of plant matter and fruit, with some species also eating dung. Octodontidae comprises fourteen extant species in seven genera.

Not assigned to a named subfamily – seven genera
Name Authority and species Range Size and ecology
Aconaemys
(rock rat)

Illustration of a rock rat

Ameghino, 1891

Three species
Chile and western Argentina
Map of range
Size range: 14–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail (Porter's rock rat)

Habitats: Rocky areas and forest

Diets: Roots, fruit, and other vegetation
Octodon
(degu)

A degu with a paw raised, viewed head-on

Bennett, 1823

Four species
Central Chile and western Argentina
Map of range
Size range: 12 cm (5 in) long, plus 16 cm (6 in) tail (Pacific degu) to 22 cm (9 in) long, plus 17 cm (7 in) tail (moon-toothed degu)

Habitats: Rocky areas, forest, and shrubland

Diets: Grass, leaves, bark, herbs, seeds, and fruit, as well as dung
Octodontomys

A degu obscured by sticks and grass

Palmer, 1903

One species
Western South America
Map of range
Size: 16–19 cm (6–7 in) long, plus 10–19 cm (4–7 in) tail

Habitats: Shrubland, grassland, and rocky areas

Diet: Acacia seed pods and cactus fruits
Octomys Thomas, 1920

One species
Western Argentina
Map of range
Size: 14–33 cm (6–13 in) long, plus 10–18 cm (4–7 in) tail

Habitats: Rocky areas

Diet: Roots, bark, and cactus
Pipanacoctomys Mares, Braun, Barquez, & Díaz, 2000

One species
Northwestern Argentina
Map of range
Size: 16–18 cm (6–7 in) long, plus 12–15 cm (5–6 in) tail

Habitats: Inland wetlands

Diet: Leaves and stems
Spalacopus

Rodent with its head sticking out of the ground

Wagler, 1832

One species
Central Chile
Map of range
Size: 11–17 cm (4–7 in) long, plus 4–6 cm (2 in) tail

Habitats: Forest, savanna and grassland

Diet: Tubers and stems of Leucocoryne and other plants
Tympanoctomys
(viscacha rat)

A rat with a hairy tail against a smooth white backdrop

Yepes, 1942

Three species
Argentina
Map of range
Size range: 11 cm (4 in) long, plus 11 cm (4 in) tail (Kirchner's viscacha rat) to 16 cm (6 in) long, plus 12 cm (5 in) tail (Chalchalero viscacha rat)

Habitats: Grassland, shrubland, shrublands, and inland wetlands

Diets: Leaves and stems

Superfamily Ctenodactyloidea

Family Ctenodactylidae

Members of the Ctenodactylidae family are called ctenodactylids or gundis. They eat a variety of vegetation. Ctenodactylidae comprises five extant species in four genera.

Not assigned to a named subfamily – four genera
Name Authority and species Range Size and ecology
Ctenodactylus
(gundi)

Brown gundi

J. E. Gray, 1830

Two species
Northwestern Africa
Map of range
Size range: 12 cm (5 in) long, plus 2 cm (0.8 in) tail (Val's gundi) to 23 cm (9 in) long, plus 5 cm (2 in) tail (common gundi)

Habitats: Shrubland and rocky areas

Diets: Vegetation
Felovia

Brown gundi

Lastaste, 1886

One species
Western Africa
Map of range
Size: 16–19 cm (6–7 in) long, plus 6–8 cm (2–3 in) tail

Habitats: Savanna, shrubland, and rocky areas

Diet: Vegetation
Massoutiera

Drawing of brown gundi

Lastaste, 1885

One species
Northwestern Africa
Map of range
Size: 12–21 cm (5–8 in) long, plus 3–9 cm (1–4 in) tail

Habitats: Rocky areas

Diet: Leaves, stems, seeds, and flowers
Pectinator

Drawing of brown gundi

Blyth, 1856

One species
Eastern Africa
Map of range
Size: 15–19 cm (6–7 in) long, plus 6–8 cm (2–3 in) tail

Habitats: Shrubland, grassland, and rocky areas

Diet: Leaves, stems, seeds, and flowers
Family Diatomyidae

Members of the Diatomyidae family are called diatomyids. The family contains a single herbivorous species.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Laonastes

Gray rock rat

Jenkins, Kilpatrick, Robinson, & Timmins, 2005

One species
Laos and Vietnam
Map of range
Size: 21–30 cm (8–12 in) long, plus 12–17 cm (5–7 in) tail

Habitats: Forest and rocky areas

Diet: Grass, seeds, and other vegetation

Family Hystricidae

Members of the Hystricidae family are called hystricids or Old World porcupines. They are primarily herbivores and eat grasses, bulbs, tubers, fruit, bark, and roots, though insects and carrion are occasionally eaten. Hystricidae comprises eleven extant species in three genera.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Atherurus
(brush-tailed porcupine)

Brown porcupine

F. Cuvier, 1829

Two species
Southeast Asia, southern China and Central Africa (outlined in blue)
Map of range
Size range: 36–60 cm (14–24 in) long, plus 10–26 cm (4–10 in) tail (African brush-tailed porcupine)

Habitats: Forest and shrubland

Diets: Vegetation, bark, roots, tubers, and fruit, as well as cultivated crops, insects, and carrion
Hystrix
(Old World porcupine)

Brown and white porcupine

Linnaeus, 1758

Eight species
Africa, Italy, and Asia (solid colors)
Map of range
Size range: 45–93 cm (18–37 in) long, plus 6–17 cm (2–7 in) tail (crested porcupine)

Habitats: Forest, grassland, shrubland, rocky areas, and savanna

Diets: Bark, roots, tubers, rhizomes, bulbs, fruit, and cultivated crops, as well as carrion, insects, and small vertebrates
Trichys

Brown porcupine under a desk

Günther, 1877

One species
Brunei, Indonesia, and Malaysia
Map of range
Size: 35–48 cm (14–19 in) long, plus 17–24 cm (7–9 in) tail

Habitats: Forest

Diet: Seeds, vegetation from trees, shrubs, and bamboo

Parvorder Phiomorpha

Family Bathyergidae

Members of the Bathyergidae family are called bathyergids, blesmols, or mole-rats. They are omnivores, and primarily eat roots, bulbs, and tubers, as well as earthworms and insects. Bathyergidae comprises twenty-one extant species in five genera.

Not assigned to a named subfamily – five genera
Name Authority and species Range Size and ecology
Bathyergus
(dune mole-rat)

Brown mole-rat

Illiger, 1811

Two species
South Africa and Namibia
Map of range
Size range: 17 cm (7 in) long, plus 4 cm (2 in) tail (Namaqua dune mole-rat) to 35 cm (14 in) long, plus 4 cm (2 in) tail (Cape dune mole-rat)

Habitats: Desert and shrubland

Diets: Bulbs and fleshy roots
Cryptomys
(common mole-rat)

Brown mole-rat

Gray, 1864

Four species
Southern Africa
Map of range
Size range: 13 cm (5 in) long, plus 1 cm (0.4 in) tail (Mahali mole-rat) to 18 cm (7 in) long, plus 3 cm (1 in) tail (Natal mole-rat)

Habitats: Grassland and shrubland

Diets: Roots, bulbs, tubers, and aloe leaves, as well as earthworms and insects
Fukomys
(tropical mole-rat)

Brown mole-rat

Kock, Ingram, Frabotta, Honeycutt, & Burda, 2006

Thirteen species
Central and southern Africa
Map of range
Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (Caroline's mole-rat) to 26 cm (10 in) long, plus 4 cm (2 in) tail (Mechow's mole-rat)

Habitats: Grassland, savanna, forest, and shrubland

Diets: Roots, bulbs, tubers, and aloe leaves, as well as earthworms and insects
Georychus

Brown mole-rat

Illiger, 1811

One species
South Africa and Lesotho
Map of range
Size: 8–23 cm (3–9 in) long, plus 1–3 cm (0.4–1.2 in) tail

Habitats: Shrubland and forest

Diet: Tubers, roots, and bulbs
Heliophobius

Brown mole-rat

Peters, 1846

One species
Southeastern Africa
Map of range
Size: 13–20 cm (5–8 in) long, plus 0.5–2 cm (0.2–0.8 in) tail

Habitats: Savanna, shrubland, grassland, and caves

Diet: Tubers and bulbs
Family Heterocephalidae

Members of the Heterocephalidae family are called heterocephalids. It includes a single species, which eats tubers, roots, and corms.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Heterocephalus

White mole rat

Rüppell, 1842

One species
Eastern Africa
Map of range
Size: 7–11 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail

Habitats: Savanna, shrubland, grassland, and caves

Diet: Tubers, roots, and corms
Family Petromuridae

Members of the Petromuridae family are called petromurids. It contains a single species, which eats vegetation, seeds, and berries.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Petromus

Gray dassie rat

A. Smith, 1831

One species
Southwestern Africa
Map of range
Size: 13–22 cm (5–9 in) long, plus 11–18 cm (4–7 in) tail

Habitats: Shrubland and rocky areas

Diet: Vegetation, seeds, and berries
Family Thryonomyidae

Members of the Thryonomyidae family are called thryonomyids or cane rats. It contains two species, which eats grass and cane, as well as bark, nuts, and fruit.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Thryonomys
(cane rat)

Brown cane rat

Fitzinger, 1867

Two species
Sub-Saharan Africa
Map of range
Size range: 41 cm (16 in) long, plus 11 cm (4 in) tail (lesser cane rat) to 77 cm (30 in) long, plus 20 cm (8 in) tail (greater cane rat)

Habitats: Shrubland, grassland, inland wetlands, and forest

Diets: Grass and cane, as well as bark, nuts, and fruit

Suborder Myomorpha

Superfamily Dipodoidea

Family Dipodidae

Members of the Dipodidae family are called dipodids or jerboas. They are omnivores, and eat a variety of vegetation as well as insects, arachnids, and lizards. Dipodidae comprises 33 extant species in 13 genera, which are divided into 4 subfamilies: Allactaginae contains 16 species, Cardiocraniinae contains 7 species, Dipodinae contains 9, and Euchoreutinae contains a single species.

Subfamily Allactaginae – Vinogradov, 1925 – four genera
Name Authority and species Range Size and ecology
Allactaga
(five-toed jerboa)

Brown jerboa

F. Cuvier, 1836

Eight species
Northern Africa, central and western Asia, and eastern Europe
Map of range
Size range: 9 cm (4 in) long, plus 16 cm (6 in) tail (Hotson's jerboa) to 23 cm (9 in) long, plus 30 cm (12 in) tail (great jerboa)

Habitats: Desert, coastal marine, shrubland, grassland, and forest

Diets: Vegetation, seeds, roots, tubers, insects, and larvae, as well as molluscs
Allactodipus Kolesnikov, 1937

One species
Central Asia
Map of range
Size: 11–14 cm (4–6 in) long, plus 17–20 cm (7–8 in) tail

Habitats: Desert

Diet: Vegetation and insects
Pygeretmus
(fat-tailed jerboa)
Gloger, 1841

Three species
Central Asia
Map of range
Size range: 7 cm (3 in) long, plus 7 cm (3 in) tail (lesser fat-tailed jerboa) to 14 cm (6 in) long, plus 18 cm (7 in) tail (dwarf fat-tailed jerboa)

Habitats: Shrubland and desert

Diets: Succulent plants and other vegetation, bulbs, rhizomes, roots, spiders, and insects
Scarturus
(four- and five-toed jerboa)

Brown jerboa

Gloger, 1841

Four species
Central and western Asia
Map of range
Size range: 7 cm (3 in) long, plus 14 cm (6 in) tail (Euphrates jerboa) to 15 cm (6 in) long, plus 26 cm (10 in) tail (Williams's jerboa)

Habitats: Shrubland, grassland, and desert

Diets: Sprouts, seeds, underground plant parts, other vegetation, and insects
Subfamily Cardiocraniinae – Vinogradov, 1925 – three genera
Name Authority and species Range Size and ecology
Cardiocranius

Brown jerboa

Satunin, 1903

One species
East-central Asia
Map of range
Size: 5–7 cm (2–3 in) long, plus 6–10 cm (2–4 in) tail

Habitats: Desert

Diet: Seeds
Salpingotulus

Brown jerboas

Pavlinov, 1980

One species
Pakistan
Map of range
Size: 4–5 cm (2 in) long, plus 7–10 cm (3–4 in) tail

Habitats: Desert

Diet: Grass seeds, stems, and other vegetation
Salpingotus
(three-toed dwarf jerboa)

Brown jerboa

Vinogradov, 1922

Five species
Central Asia
Map of range
Size range: 4 cm (2 in) long, plus 8 cm (3 in) tail (Heptner's pygmy jerboa) to 7 cm (3 in) long, plus 14 cm (6 in) tail (Kozlov's pygmy jerboa)

Habitats: Shrubland, desert, and grassland

Diets: Insects, arachnids, and vegetation
Subfamily Dipodinae – G. Fischer, 1817 – five genera
Name Authority and species Range Size and ecology
Dipus

Brown jerboa

Zimmermann, 1780

One species
Western, central, and eastern Asia Size: 10–15 cm (4–6 in) long, plus 12–20 cm (5–8 in) tail

Habitats: Shrubland and desert

Diet: All parts of plants, as well as insects
Eremodipus Vinogradov, 1930

One species
Central Asia Size: 10–12 cm (4–5 in) long, plus 13–17 cm (5–7 in) tail

Habitats: Desert

Diet: Roots, sprouts, seeds, grains, and vegetables
Jaculus
(desert jerboa)

Brown jerboa

Erxleben, 1777

Three species
Northern Africa, Middle East, and central Asia Size range: 9 cm (4 in) long, plus 17 cm (7 in) tail (greater Egyptian jerboa) to 15 cm (6 in) long, plus 22 cm (9 in) tail (Blanford's jerboa)

Habitats: Desert, coastal marine, shrubland, grassland, and rocky areas

Diets: Roots, sprouts, seeds, grains, and vegetables
Paradipus Vinogradov, 1930

One species
Central Asia Size: 14–17 cm (6–7 in) long, plus 18–22 cm (7–9 in) tail

Habitats: Desert

Diet: All parts of desert plants
Stylodipus
(thick-tailed three-toed jerboa)

Brown jerboa

Allen, 1925

Three species
Ukraine and western and central Asia Size range: 11 cm (4 in) long, plus 13 cm (5 in) tail (thick-tailed three-toed jerboa) to 14 cm (6 in) long, plus 17 cm (7 in) tail (Andrews's three-toed jerboa)

Habitats: Shrubland, desert, grassland, and forest

Diets: Lichen, rhizomes, bulbs, seeds, and wheat
Subfamily Euchoreutinae – Lyon, 1901 – one genus
Name Authority and species Range Size and ecology
Euchoreutes

Drawing of brown jerboa

W. L. Sclater, 1891

One species
Southern Mongolia and China Size: 9–11 cm (4 in) long, plus 14–18 cm (6–7 in) tail

Habitats: Shrubland and desert

Diet: Insects and lizards
Family Sminthidae

Members of the Sminthidae family are called sminthids or birch mice. They are omnivores and primarily eat seeds, berries, and insects. Sminthidae comprises sixteen extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Sicista
(birch mouse)

Brown birch mouse

Gray, 1827

Sixteen species
Asia and Europe
Map of range
Size range: 4 cm (2 in) long, plus 8 cm (3 in) tail (Caucasian birch mouse) to 8 cm (3 in) long, plus 11 cm (4 in) tail (Altai birch mouse)

Habitats: Inland wetlands, shrubland, forest, and grassland

Diets: Seeds, berries, and insects
Family Zapodidae

Members of the Zapodidae family are called zapodids or jumping mice. They are omnivores and eat a variety of vegetation and insects. Zapodidae comprises five extant species in three genera.

Not assigned to a named subfamily – three genera
Name Authority and species Range Size and ecology
Eozapus Preble, 1899

One species
China
Map of range
Size: 7–10 cm (3–4 in) long, plus 11–15 cm (4–6 in) tail

Habitats: Forest, shrubland, and grassland

Diet: Vegetation and insects
Napaeozapus

Brown mouse

Preble, 1899

One species
Eastern Canada and eastern United States
Map of range
Size: 8–10 cm (3–4 in) long, plus 11–16 cm (4–6 in) tail

Habitats: Forest, shrubland, and inland wetlands

Diet: Seeds, fungi, and insects, as well as fruit, nuts, and other vegetation
Zapus
(meadow jumping mouse)

Brown mouse

Coues, 1875

Three species
Canada and United States
Map of range
Size range: 7 cm (3 in) long, plus 10 cm (4 in) tail (meadow jumping mouse) to 11 cm (4 in) long, plus 16 cm (6 in) tail (Pacific jumping mouse)

Habitats: Forest, shrubland, grassland, and inland wetlands

Diets: Seeds, as well as fungi, nuts, berries, fruit, and insects

Superfamily Muroidea

Family Calomyscidae

Members of the Calomyscidae family are called calomyscids or brush-tailed mice. They are omnivores and primarily eat seeds, as well as flowers, leaves, and animal matter. Calomyscidae comprises eight extant species in a single genus.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Calomyscus
(mouse-like hamster)

Brown hamster

Thomas, 1905

Central Asia
Map of range
Size range: 6 cm (2 in) long, plus 5 cm (2 in) tail (Tsolov's mouse-like hamster) to 10 cm (4 in) long, plus 11 cm (4 in) tail (great Balkhan mouse-like hamster)

Habitats: Shrubland, grassland and rocky areas

Diets: Seeds, as well as flowers, leaves, and animal matter
Family Cricetidae

Members of the Cricetidae family are called cricetids and include hamsters, voles, lemmings, muskrats, and New World rats and mice. They eat vegetation, seeds, and insects, and some also eat fungi, fish, and small animals. Cricetidae comprises 694 extant species, the second-most of any mammal family, divided into 143 genera. These genera are divided into five subfamilies: Arvicolinae, containing 158 species of voles, lemmings, and muskrats; Cricetinae, containing 18 species of hamsters; Neotominae, containing 123 species of deer mice, packrats, and grasshopper mice; Sigmodontinae, containing 385 species of New World rats and mice; and Tylomyinae, containing 10 species of vesper and climbing rats. Additionally, one species in Neotominae and ten species in Sigmodontinae were driven extinct after 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s.

Subfamily Arvicolinae – Gray, 1821 – 30 genera
Name Authority and species Range Size and ecology
Alexandromys
(grass vole)

Gray vole

Ognev, 1914

Eleven species
Eastern and northern Asia, Europe, and northwestern North America Size range: 7 cm (3 in) long, plus 2 cm (0.8 in) tail (tundra vole) to 18 cm (7 in) long, plus 8 cm (3 in) tail (reed vole)

Habitats: Forest, shrubland, inland wetlands, desert, rocky areas, and grassland

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Alticola
(high mountain vole)

Gray and brown vole

Blanford, 1881

Twelve species
Asia Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (lemming vole) to 14 cm (6 in) long, plus 5 cm (2 in) tail (flat-headed vole)

Habitats: Forest, rocky areas, shrubland, and grassland

Diets: Stems and leaves
Arborimus
(tree vole)

Brown vole

Taylor, 1915

Three species
Western United States Size range: 9–11 cm (4 in) long, plus 6–9 cm (2–4 in) tail (red tree vole)

Habitats: Forest and inland wetlands

Diets: Conifer needles
Arvicola
(water vole)

Brown vole

Lacépède, 1799

Three species
Europe and Asia Size range: 12–23 cm (5–9 in) long, plus 5–14 cm (2–6 in) tail (European water vole)

Habitats: Forest, inland wetlands, and grassland

Diets: Aquatic plants, herbs, grass, twigs, buds, roots, bulbs, and fruit
Caryomys
(brownish vole)
Thomas, 1911

Two species
Central China Size range: 8–11 cm (3–4 in) long, plus 4–6 cm (2 in) tail (Ganzu vole)

Habitats: Forest

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Chionomys
(snow vole)

Brown vole

Miller, 1908

Three species
Europe and western Asia Size range: 10 cm (4 in) long, plus 4 cm (2 in) tail (European snow vole) to 16 cm (6 in) long, plus 11 cm (4 in) tail (Robert's snow vole)

Habitats: Forest and rocky areas

Diets: Grass, herbs, and seeds
Clethrionomys
(bank vole)

Brown vole

Gistel, 1850

Five species
Europe, Asia, and North America Size range: 8–14 cm (3–6 in) long, plus 3–8 cm (1–3 in) tail (bank vole)

Habitats: Forest, rocky areas, shrubland, and grassland

Diets: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects
Craseomys
(grey-sided vole)

Brown vole

Miller, 1900

Six species
Northern Europe and northern and eastern Asia Size range: 7 cm (3 in) long, plus 4 cm (2 in) tail (Japanese red-backed vole) to 15 cm (6 in) long, plus 7 cm (3 in) tail (grey red-backed vole)

Habitats: Forest, shrubland, inland wetlands, rocky areas, and grassland

Diets: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects
Dicrostonyx
(collared lemming)

Gray and brown lemming

Gloger, 1841

Eight species
Russia and northern North America Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (Arctic lemming) to 16 cm (6 in) long, plus 3 cm (1 in) tail (northern collared lemming)

Habitats: Rocky areas, shrubland, inland wetlands, and grassland

Diets: Fruit, flowers, grass, sedges, buds, twigs, and bark
Dinaromys Kretzoi, 1955

One species
Southern Europe
Map of range
Size: 13–16 cm (5–6 in) long, plus 8–11 cm (3–4 in) tail

Habitats: Rocky areas

Diet: Grass
Ellobius
(mole vole)

Brown vole

Fischer von Waldheim, 1814

Five species
Eastern Europe and western and central Asia Size range: 8 cm (3 in) long, plus 0.5 cm (0 in) tail (northern mole vole) to 15 cm (6 in) long, plus 2 cm (0.8 in) tail (southern mole vole)

Habitats: Desert, shrubland, and grassland

Diets: Bulbs, tubers, and other underground plant parts
Eolagurus
(steppe lemming)

Drawing of brown lemmings

Argiropulo, 1946

Two species
Central Asia Size range: 12 cm (5 in) long, plus 0.5 cm (0 in) tail (Przewalski's steppe lemming) to 16 cm (6 in) long, plus 3 cm (1 in) tail (yellow steppe lemming)

Habitats: Shrubland and grassland

Diets: Bulbs, seeds, and roots
Eothenomys
(Père David's vole)

Drawing of brown vole

Miller, 1896

Eight species
Eastern Asia Size range: 7 cm (3 in) long, plus 2 cm (0.8 in) tail (Père David's vole) to 14 cm (6 in) long, plus 7 cm (3 in) tail (Kachin red-backed vole)

Habitats: Forest, rocky areas, shrubland, and grassland

Diets: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects
Hyperacrius
(Kashmir vole)
Miller, 1896

Two species
Northern Pakistan and western China Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (True's vole) to 14 cm (6 in) long, plus 5 cm (2 in) tail (Murree vole)

Habitats: Forest, shrubland, and grassland

Diets: Grass, stems, and roots
Lagurus

Gray lemming

Gloger, 1841

One species
Eastern Europe and western, central, and eastern Asia Size: 7–13 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail

Habitats: Shrubland and grassland

Diet: Green vegetation, tubers, and bulbs
Lasiopodomys
(narrow-headed steppe vole)

Brown vole

Lataste, 1887

Two species
Eastern Asia Size range: 9 cm (4 in) long, plus 2 cm (0.8 in) tail (Mandarin vole) to 15 cm (6 in) long, plus 3 cm (1 in) tail (Brandt's vole)

Habitats: Shrubland and grassland

Diets: Grass, vegetation, and roots
Lemmiscus

Gray vole

Thomas, 1912

One species
Western United States and western Canada
Map of range
Size: 9–12 cm (4–5 in) long, plus 1–3 cm (0–1 in) tail

Habitats: Shrubland and grassland

Diet: Green vegetation
Lemmus
(lemming)

Brown and yellow lemming

Link, 1795

Five species
Northern Europe, northern Asia, and western North America Size range: 8 cm (3 in) long, plus 1 cm (0.4 in) tail (Amur lemming) to 16 cm (6 in) long, plus 2 cm (0.8 in) tail (West Siberian lemming)

Habitats: Forest, shrubland, inland wetlands, and grassland

Diets: Sedges, grass, bark, leaves, berries, lichens, and roots
Microtus
(meadow vole)

Brown vole

Schrank, 1798

57 species
North America, Central America, Europe, and Asia Size range: 6 cm (2 in) long, plus 1 cm (0.4 in) tail (Afghan vole) to 18 cm (7 in) long, plus 10 cm (4 in) tail (North American water vole)

Habitats: Forest, intertidal marine, coastal marine, savanna, shrubland, desert, inland wetlands, rocky areas, and grassland

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Myopus

Gray lemming

Miller, 1910

One species
Northern Europe and northern Asia Size: 9–12 cm (4–5 in) long, plus 1–2 cm (0.4–0.8 in) tail

Habitats: Forest and inland wetlands

Diet: Moss, stems, and bark
Neodon
(mountain vole)

Brown vole

Horsfield, 1841

Six species
China and southeastern Asia Size range: 8 cm (3 in) long, plus 2 cm (0.8 in) tail (Chinese scrub vole) to 15 cm (6 in) long, plus 5 cm (2 in) tail (plateau vole)

Habitats: Forest, shrubland, rocky areas, and grassland

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Neofiber

Brown muskrat

True, 1884

One species
Southeastern United States
Map of range
Size: 18–21 cm (7–8 in) long, plus 9–17 cm (4–7 in) tail

Habitats: Grassland and inland wetlands

Diet: Maidencane, as well as rushes, sedges, sawgrass, and mangrove bark
Ondatra

Brown muskrat

Link, 1795

One species
North America, Europe, and Asia
Map of range
Size: 27–30 cm (11–12 in) long, plus 20–25 cm (8–10 in) tail

Habitats: Inland wetlands

Diet: Cattails, bulrushes, grass, and other vegetation, as well as crabs, crayfish, mussels, and small fish
Phaiomys

Drawing of brown vole

Blyth, 1863

One species
Western China Size: 9–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail

Habitats: Forest, grassland, rocky areas, and shrubland

Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Phenacomys
(heather vole)

Drawing of brown voles

Merriam, 1889

Two species
Northwestern United States and Canada Size range: 8–12 cm (3–5 in) long, plus 2–4 cm (0.8–1.6 in) tail (eastern heather vole)

Habitats: Forest, shrubland, and grassland

Diets: Bark, buds, heaths, forbs, berries, and seeds
Proedromys
(Duke of Bedford's vole)
Thomas, 1911

Two species
Central China Size range: 9 cm (4 in) long, plus 3 cm (1 in) tail (Duke of Bedford's vole) to 14 cm (6 in) long, plus 9 cm (4 in) tail (Liangshan vole)

Habitats: Forest and grassland

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Prometheomys Satunin, 1901

One species
Western Asia Size: 12–17 cm (5–7 in) long, plus 3–6 cm (1–2 in) tail

Habitats: Grassland

Diet: Green vegetation and roots
Stenocranius
(narrow-headed vole)

Drawing of brown vole

Kaschtschenko, 1901

Two species
Northern Asia Size range: 8–15 cm (3–6 in) long, plus 1–5 cm (0.4–2.0 in) tail (narrow-headed vole)

Habitats: Grassland

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Synaptomys
(bog lemming)

Brown lemming

Baird, 1857

Two species
Canada and northern United States Size range: 9 cm (4 in) long, plus 1 cm (0.4 in) tail (northern bog lemming) to 14 cm (6 in) long, plus 3 cm (1 in) tail (southern bog lemming)

Habitats: Forest, shrubland, inland wetlands, and grassland

Diets: Green vegetation as well as slugs, snails, and other invertebrates
Volemys
(Szechuan vole)
Zagorodnyuk, 1990

Two species
Central China Size range: 8 cm (3 in) long, plus 4 cm (2 in) tail (Szechuan vole) to 13 cm (5 in) long, plus 8 cm (3 in) tail (Marie's vole)

Habitats: Forest, rocky areas, and grassland

Diets: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation
Subfamily Cricetinae – Waldheim, 1817 – seven genera
Name Authority and species Range Size and ecology
Allocricetulus
(Mongolian hamster)
Argiropulo, 1932

Two species
East-central and central Asia Size range: 10 cm (4 in) long, plus 1 cm (0.4 in) tail (Mongolian hamster) to 14 cm (6 in) long, plus 4 cm (2 in) tail (Eversmann's hamster)

Habitats: Grassland and shrubland

Diets: Grains, beans, lentils, roots, and plant material, as well as insects and frogs
Cansumys Allen, 1928

One species
Central China Size: 12–14 cm (5–6 in) long, plus 9–11 cm (4 in) tail

Habitats: Forest

Diet: Leaves and grasses
Cricetulus
(ratlike hamster)

Brown hamster

A. Milne-Edwards, 1867

Three species
Central and eastern Asia Size range: 7 cm (3 in) long, plus 1 cm (0.4 in) tail (Chinese striped hamster) to 14 cm (6 in) long, plus 5 cm (2 in) tail (long-tailed dwarf hamster)

Habitats: Grassland, forest, inland wetlands, desert, rocky areas, and shrubland

Diets: Shoots, seeds, and insects
Cricetus
(golden hamster)

Brown hamster

Leske, 1779

One species
Europe and western and central Asia
Map of range
Size: 16–32 cm (6–13 in) long, plus 2–7 cm (1–3 in) tail

Habitats: Shrubland and grassland

Diet: Grains, beans, lentils, roots, and plant material, as well as insects and frogs
Mesocricetus

Brown hamster

Nehring, 1898

Four species
Western Asia and eastern Europe Size range: 12 cm (5 in) long, plus 1 cm (0.4 in) tail (golden hamster) to 22 cm (9 in) long, plus 2 cm (0.8 in) tail (Ciscaucasian hamster)

Habitats: Grassland and shrubland

Diets: Omnivorous, including a variety of vegetation, seeds, fruit, insects, and small vertebrates
Nothocricetulus

Gray hamster

Lebedev, Bannikova, Neumann, Ushakova, Ivanova, & Surov, 2018

One species
Asia and eastern Europe Size range: 8–15 cm (3–6 in) long, plus 1–4 cm (0.4–1.6 in)

Habitats: Forest, shrubland, grassland, and desert

Diets: Shoots, seeds, and insects
Phodopus
(dwarf hamster)

Brown hamster

Miller, 1910

Three species
East-central Asia and Kazakhstan and southern Russia Size range: 7 cm (3 in) long, plus 0.5 cm (0.2 in) tail (Roborovski dwarf hamster) to 11 cm (4 in) long, plus 2 cm (0.8 in) tail (Campbell's dwarf hamster)

Habitats: Desert and shrubland

Diets: Seeds and plant material
Tscherskia

Brown hamster

Ogniov, 1914

One species
Eastern Asia Size: 14–22 cm (6–9 in) long, plus 6–11 cm (2–4 in) tail

Habitats: Inland wetlands

Diet: Seeds as well as shoots
Urocricetus Satunin, 1903

Two species
China Size: 8 cm (3 in) long, plus 2 cm (1 in) tail (Tibetan dwarf hamster) to 12 cm (5 in) long, plus 7 cm (3 in) tail (Kam dwarf hamster)

Habitats: Forest, shrubland, grassland, and inland wetlands

Diet: Shoots, seeds, and insects
Subfamily Neotominae – Merriam, 1894 – sixteen genera
Name Authority and species Range Size and ecology
Baiomys
(New World pygmy mouse)

Gray mouse

True, 1894

Two species
Southern United States, Mexico, and northern Central America Size range: 5 cm (2 in) long, plus 3 cm (1 in) tail (northern pygmy mouse) to 8 cm (3 in) long, plus 6 cm (2 in) tail (Mexican pygmy mouse)

Habitats: Grassland, desert, forest, and shrubland

Diets: Cactus stems and fruit, green vegetation, seeds, grass, and insects
Habromys
(slender-tailed deer mouse)
Hooper & Musser, 1964

Six species
Mexico and northern Central America Size range: 7 cm (3 in) long, plus 7 cm (3 in) tail (delicate deer mouse) to 14 cm (6 in) long, plus 15 cm (6 in) tail (Ixtlán deer mouse)

Habitats: Forest

Diets: Plant material and invertebrates
Hodomys

Mouse jawbone

Merriam, 1894

One species
Southern Mexico Size: 16–22 cm (6–9 in) long, plus 14–22 cm (6–9 in) tail

Habitats: Shrubland

Diet: Plant material and invertebrates
Isthmomys Hooper & Musser, 1964

Two species
Panama and northwestern Colombia Size range: 15–18 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail (Mount Pirri isthmus rat)

Habitats: Forest

Diets: Plant material and invertebrates
Megadontomys
(giant deer mouse)

Brown mouse

Merriam, 1898

Three species
Southern Mexico Size range: 13 cm (5 in) long, plus 17 cm (7 in) tail (Nelson's giant deer mouse) to 17 cm (7 in) long, plus 19 cm (7 in) tail (Thomas's giant deer mouse)

Habitats: Forest

Diets: Berries and seeds
Nelsonia
(diminutive woodrat)
Merriam, 1897

Two species
Central Mexico Size range: 11 cm (4 in) long, plus 10 cm (4 in) tail (diminutive woodrat) to 14 cm (6 in) long, plus 13 cm (5 in) tail (Goldman's diminutive woodrat)

Habitats: Forest

Diets: Conifer needles
Neotoma
(pack rat)

Gray mouse

Say & Ord, 1825

20 species
North America and Central America Size range: 14 cm (6 in) long, plus 10 cm (4 in) tail (Arizona woodrat) to 25 cm (10 in) long, plus 21 cm (8 in) tail (big-eared woodrat)

Habitats: Desert, shrubland, grassland, rocky areas, forest, caves, coastal marine, and inland wetlands

Diets: Roots, stems, and leaves, as well as seeds and invertebrates
Neotomodon

Brown mouse

Merriam, 1898

One species
Central Mexico Size: 9–13 cm (4–5 in) long, plus 7–11 cm (3–4 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Ochrotomys

Drawing of brown mouse

Osgood, 1909

One species
Eastern United States Size: 5–12 cm (2–5 in) long, plus 5–10 cm (2–4 in) tail

Habitats: Shrubland and forest

Diet: Seeds
Onychomys
(grasshopper mouse)

Brown and white mouse

Baird, 1857

Three species
United States and Mexico Size range: 8 cm (3 in) long, plus 4 cm (2 in) tail (Mearns's grasshopper mouse) to 13 cm (5 in) long, plus 7 cm (3 in) tail (northern grasshopper mouse)

Habitats: Grassland, desert, and shrubland

Diets: Insects, scorpions, and small vertebrates, as well as plant material
Osgoodomys

Brown mouse

Hooper & Musser, 1964

One species
Southwestern Mexico Size: 10–15 cm (4–6 in) long, plus 9–13 cm (4–5 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Peromyscus
(deer mouse)

Brown mouse

Gloger, 1841

57 species
North America and Central America Size range: 7 cm (3 in) long, plus 8 cm (3 in) tail (cactus mouse) to 15 cm (6 in) long, plus 16 cm (6 in) tail (brown deer mouse)

Habitats: Desert, shrubland, grassland, rocky areas, forest, caves, intertidal marine, coastal marine, and inland wetlands

Diets: A wide variety of plant material, fungi, insects, other invertebrates, and carrion, as well as worms, molluscs, and small vertebrates
Podomys

Gray mouse

Osgood, 1909

One species
Southeastern United States Size: 9–12 cm (4–5 in) long, plus 8–11 cm (3–4 in) tail

Habitats: Savanna, forest, grassland, and shrubland

Diet: Seeds, nuts, fungi, vegetation, insects, and other invertebrates
Reithrodontomys
(harvest mouse)

Brown mouse

Giglioli, 1873

21 species
North America and Central America Size range: 5 cm (2 in) long, plus 5 cm (2 in) tail (plains harvest mouse) to 13 cm (5 in) long, plus 13 cm (5 in) tail (narrow-nosed harvest mouse)

Habitats: Desert, shrubland, grassland, forest, intertidal marine, and inland wetlands

Diets: Seeds and shoots, as well as insects
Scotinomys
(singing mouse)

Drawing of brown mouse

Thomas, 1913

Two species
Southern Mexico and Central America Size range: 6 cm (2 in) long, plus 4 cm (2 in) tail (Alston's brown mouse) to 9 cm (4 in) long, plus 8 cm (3 in) tail (Chiriqui brown mouse)

Habitats: Grassland and forest

Diets: Insects
Xenomys

Gray mouse

Merriam, 1892

One species
Southwestern Mexico Size: 15–17 cm (6–7 in) long, plus 14–17 cm (6–7 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Subfamily Sigmodontinae – Wagner, 1843 – 87 genera
Name Authority and species Range Size and ecology
Abrawayaomys Cunha & Cruz, 1979

One species
Southeastern Brazil Size: 8–14 cm (3–6 in) long, plus 8–15 cm (3–6 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Abrothrix
(soft-haired mouse)

Brown and white mouse

Waterhouse, 1837

Eight species
Western South America Size range: 6 cm (2 in) long, plus 4 cm (2 in) tail (Andean Altiplano mouse) to 14 cm (6 in) long, plus 10 cm (4 in) tail (long-haired grass mouse)

Habitats: Rocky areas, inland wetlands, grassland, forest, and shrubland

Diets: Insects, berries, seeds, and fungi
Aegialomys
(Galápagos rice rat)

Gray rat

Weksler, Percequillo, & Voss, 2006

Two species
Ecuador and western Peru Size range: 10–16 cm (4–6 in) long, plus 12–18 cm (5–7 in) tail (Galápagos rice rat)

Habitats: Desert, forest, and shrubland

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Aepeomys
(montane mouse)
Thomas, 1898

Two species
Western Venezuela Size range: 10–13 cm (4–5 in) long, plus 11–15 cm (4–6 in) tail (Reig's montane mouse)

Habitats: Grassland, forest, and shrubland

Diets: Vegetation, seeds, and insects
Akodon
(grass mouse)

Brown mouse

Meyen, 1833

39 species
South America Size range: 6 cm (2 in) long, plus 4 cm (2 in) tail (Caparaó grass mouse) to 16 cm (6 in) long, plus 11 cm (4 in) tail (variable grass mouse)

Habitats: Rocky areas, inland wetlands, savanna, grassland, forest, and shrubland

Diets: Plants and invertebrates
Amphinectomys Malygin, 1994

One species
Northern Peru Size: 18–19 cm (7 in) long, plus 17–21 cm (7–8 in) tail

Habitats: Inland wetlands and forest

Diet: Unknown
Andalgalomys
(chaco mouse)
Williams & Mares, 1978

Two species
Southern Bolivia, Paraguay, and Northern Argentina Size range: 7 cm (3 in) long, plus 8 cm (3 in) tail (Olrog's chaco mouse) to 13 cm (5 in) long, plus 14 cm (6 in) tail (Pearson's chaco mouse)

Habitats: Grassland and shrubland

Diets: Grass, grain, and mesquite fruit
Andinomys Thomas, 1902

One species
Western South America Size: 13–19 cm (5–7 in) long, plus 10–16 cm (4–6 in) tail

Habitats: Grassland and shrubland

Diet: Vegetation
Anotomys , Genus Anotomys

One species
Ecuador and Colombia Size: 10–13 cm (4–5 in) long, plus 12–16 cm (5–6 in) tail

Habitats: Inland wetlands, grassland, and forest

Diet: Fish
Auliscomys
(big-eared mouse)

Brown mouse

Thomas, 1906

Three species
Western South America Size range: 9 cm (4 in) long, plus 4 cm (2 in) tail (Andean big-eared mouse) to 14 cm (6 in) long, plus 11 cm (4 in) tail (painted big-eared mouse)

Habitats: Rocky areas, grassland, inland wetlands, and shrubland

Diets: Seeds, vegetation, and lichen
Bibimys
(crimson-nosed rat)

Brown mouse

Massoia, 1979

Three species
Paraguay, Argentina, and southern Brazil Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (large-lipped crimson-nosed rat) to 11 cm (4 in) long, plus 8 cm (3 in) tail (Torres's crimson-nosed rat)

Habitats: Grassland, forest, inland wetlands, and shrubland

Diets: Grass and seeds
Blarinomys Thomas, 1896

One species
Northeastern Argentina and southern Brazil Size: 9–13 cm (4–5 in) long, plus 3–6 cm (1–2 in) tail

Habitats: Forest

Diet: Insects and worms
Brucepattersonius
(brucie)
Hershkovitz, 1998

Seven species
Northeastern Argentina and southern Brazil Size range: 8 cm (3 in) long, plus 8 cm (3 in) tail (Arroyo of Paradise brucie) to 13 cm (5 in) long, plus 10 cm (4 in) tail (red-bellied akodont)

Habitats: Rocky areas and forest

Diets: Vegetation, seeds, and insects
Calomys
(vesper mouse)

Brown mouse

Waterhouse, 1837

Thirteen species
South America Size range: 5 cm (2 in) long, plus 4 cm (2 in) tail (Hummelinck's vesper mouse) to 17 cm (7 in) long, plus 11 cm (4 in) tail (Córdoba vesper mouse)

Habitats: Rocky areas, savanna, grassland, forest, and shrubland

Diets: Vegetation and insects
Casiomys
(Casio rice rat)

Brown rat

Voss, 2024

Six species
Mexico, Central America, and northeastern South America Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (black-eared rice rat) to 16 cm (6 in) long, plus 12 cm (5 in) tail (Chapman's rice rat)

Habitats: Forest

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Cerradomys
(Cerrado rice rat)
Weksler, Percequillo, & Voss, 2006

Four species
South America Size range: 12–19 cm (5–7 in) long, plus 14–20 cm (6–8 in) tail (Lindbergh's rice rat)

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland

Diets: Vegetation, seeds, and insects
Chelemys
(long-clawed mouse)

Drawing of brown and white mice

Thomas, 1903

Two species
Chile and southwestern Argentina Size range: 10–15 cm (4–6 in) long, plus 4–7 cm (2–3 in) tail (Andean long-clawed mouse)

Habitats: Grassland, forest, and shrubland

Diets: Arthropods, vegetation, and fungi
Chibchanomys
(water mouse)
Voss, 1988

Two species
Northwestern South America Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (Las Cajas water mouse) to 13 cm (5 in) long, plus 14 cm (6 in) tail (Chibchan water mouse)

Habitats: Grassland, forest, and inland wetlands

Diets: Aquatic invertebrates and small animals
Chilomys Thomas, 1897

One species
Northwestern South America Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail

Habitats: Forest

Diet: Omnivorous
Chinchillula Thomas, 1898

One species
Western South America
Map of range
Size: 15–17 cm (6–7 in) long, plus 9–11 cm (4 in) tail

Habitats: Shrubland and rocky areas

Diet: Vegetation
Delomys
(Atlantic Forest rat)

Brown rat

Thomas, 1917

Three species
Southeastern South America Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (montane Atlantic Forest rat) to 15 cm (6 in) long, plus 13 cm (5 in) tail (pallid Atlantic Forest rat)

Habitats: Grassland and forest

Diets: Vegetation, seeds, and insects
Deltamys

Brown mouse

Thomas, 1917

One species
Southeastern South America Size: About 10 cm (4 in)

Habitats: Inland wetlands and grassland

Diet: Plants and invertebrates
Drymoreomys

Brown rat

Percequillo, Weksler, & Costa, 2011

One species
Southeastern Brazil
Map of range
Size: 11–15 cm (4–6 in) long, plus 14–18 cm (6–7 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Eligmodontia
(gerbil mouse)
F. Cuvier, 1837

Four species
Western South America Size range: 6–10 cm (2–4 in) long, plus 5–10 cm (2–4 in) tail (Andean gerbil mouse)

Habitats: Desert, grassland, and shrubland

Diets: Grain, vegetation, and insects
Eremoryzomys Weksler, Percequillo, & Voss, 2006

One species
Northern Peru and southern Ecuador
Map of range
Size: 14–18 cm (6–7 in) long, plus 16–21 cm (6–8 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Euneomys
(chinchilla mouse)

Brown mouse

Coues, 1874

Four species
Chile and Argentina Size range: 7–16 cm (3–6 in) long, plus 4–9 cm (2–4 in) tail (Patagonian chinchilla mouse)

Habitats: Rocky areas, grassland, forest, and shrubland

Diets: Vegetation
Euryoryzomys
(broad rice rat)

Brown rat

Weksler, Percequillo, & Voss, 2006

Six species
South America Size range: 9–18 cm (4–7 in) long, plus 9–19 cm (4–7 in) tail (Russet rice rat)

Habitats: Savanna and forest

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Galenomys Thomas, 1916

One species
Western South America Size: 10–14 cm (4–6 in) long, plus 3–5 cm (1–2 in) tail

Habitats: Grassland

Diet: Seeds, vegetation, and lichen
Geoxus
(long-clawed mole mouse)

Brown and white mouse

Thomas, 1919

Two species
Chile and southern Argentina Size range: 9 cm (4 in) long, plus 3 cm (1 in) tail (long-clawed mole mouse) to 13 cm (5 in) long, plus 9 cm (4 in) tail (Pearson's long-clawed akodont)

Habitats: Forest and shrubland

Diets: Worms, insects, and other arthropods
Graomys
(gray leaf-eared mouse)

Gray mouse

Thomas, 1916

Four species
Central and southern South America Size range: 10 cm (4 in) long, plus 14 cm (6 in) tail (pale leaf-eared mouse) to 17 cm (7 in) long, plus 18 cm (7 in) tail (gray leaf-eared mouse)

Habitats: Grassland, forest, and shrubland

Diets: Grass, grain, and mesquite fruit
Gyldenstolpia

Drawing of gray rat

Pardiñas, D'Elía, & Teta, 2009

One species
Southern Paraguay and northern Argentina Size: About 22 cm (9 in)

Habitats: Inland wetlands and savanna

Diet: Vegetation, seeds, and insects
Handleyomys
(Handley's rice rat)
Voss, Gómez-Laverde, & Pacheco, 2002

Two species
Northwestern Colombia Size range: 8 cm (3 in) long, plus 7 cm (3 in) tail (Colombian rice rat) to 13 cm (5 in) long, plus 11 cm (4 in) tail (dusky-footed Handley's mouse)

Habitats: Forest

Diets: Vegetation, seeds, and insects
Holochilus
(marsh rat)

Brown rat

Brandt, 1835

Three species
South America Size range: 13 cm (5 in) long, plus 12 cm (5 in) tail (Amazonian marsh rat) to 24 cm (9 in) long, plus 24 cm (9 in) tail (Brazilian marsh rat)

Habitats: Grassland, forest, inland wetlands, and shrubland

Diets: Marsh plants and molluscs
Hylaeamys
(forest rice rat)

Brown rat

Weksler, Percequillo, & Voss, 2006

Eight species
South America Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (Azara's broad-headed oryzomys) to 18 cm (7 in) long, plus 16 cm (6 in) tail (Atlantic Forest rice rat)

Habitats: Inland wetlands, savanna, grassland, and forest

Diets: Vegetation, seeds, and insects
Ichthyomys
(crab-eating rat)

Drawing of brown and white rat

Thomas, 1893

Four species
Northwestern South America Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Pittier's crab-eating rat) to 20 cm (8 in) long, plus 16 cm (6 in) tail (Tweedy's crab-eating rat)

Habitats: Forest and inland wetlands

Diets: Fish, crabs, and aquatic insects
Irenomys Thomas, 1919

One species
Southern Chile and southwestern Argentina Size: 10–13 cm (4–5 in) long, plus 16–20 cm (6–8 in) tail

Habitats: Shrubland and forest

Diet: Fruit, seeds, and vegetation
Juliomys
(Atlantic Forest tree mouse)

Brown mouse

González, 2000

Two species
Southern Brazil and northeastern Argentina Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (montane red-rumped tree mouse) to 11 cm (4 in) long, plus 12 cm (5 in) tail (lesser Wilfred's mouse)

Habitats: Forest

Diets: Vegetation, seeds, and insects
Juscelinomys
(Brazilian burrowing mouse)

Brown mouse

Oliveira, 1965

Two species
Northeastern Bolivia and Brazil Size range: 13–20 cm (5–8 in) long, plus 8–10 cm (3–4 in) tail (Huanchaca mouse)

Habitats: Savanna, and grassland

Diets: Vegetation and insects
Kunsia

Gray rat

Hershkovitz, 1966

One species
Northern Bolivia and western Brazil Size: 18–29 cm (7–11 in) long, plus 14–20 cm (6–8 in) tail

Habitats: Grassland

Diet: Roots and grass
Lenoxus Thomas, 1909

One species
Southern Peru and western Bolivia Size: 11–17 cm (4–7 in) long, plus 13–19 cm (5–7 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Loxodontomys
(big-eared mouse)

Drawing of brown mouse

Osgood, 1947

Two species
Chile and southwestern Argentina Size range: 9–15 cm (4–6 in) long, plus 8–12 cm (3–5 in) tail (Pikumche pericote)

Habitats: Shrubland

Diets: Vegetation, fungi, and flowers
Lundomys Voss & Carleton, 1993

One species
Uruguay and southern Brazil Size: 17–24 cm (7–9 in) long, plus 19–29 cm (7–11 in) tail

Habitats: Inland wetlands, forest, and grassland

Diet: Vegetation
Megalomys †
(pilorie)

Brown mouse

Trouessart, 1881

Two species
Martinique and St. Lucia Size range: Unknown

Habitats: Unknown

Diets: Unknown
Megaoryzomys †

Rat skull pieces

Lenglet & Coppois, 1979

One species
Santa Cruz island of the Galápagos Islands Size: Unknown

Habitats: Shrubland

Diet: Unknown
Melanomys
(dark rice rat)

Brown rat

Thomas, 1902

Three species
Central America and northwestern South America Size range: 10–14 cm (4–6 in) long, plus 7–12 cm (3–5 in) tail (dusky rice rat)

Habitats: Desert, forest, and shrubland

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Microakodontomys

Brown rats

Hershkovitz, 1993

One species
Central Brazil Size: About 7 cm (3 in)

Habitats: Shrubland and grassland

Diet: Vegetation, seeds, and insects
Microryzomys
(colilargo)
Thomas, 1917

Two species
Northwestern South America Size range: 6–10 cm (2–4 in) long, plus 11–13 cm (4–5 in) tail (montane colilargo)

Habitats: Rocky areas, grassland, and forest

Diets: Seeds and vegetation
Mindomys Weksler, Percequillo, & Voss, 2006

One species
Ecuador
Map of range
Size: 17–29 cm (7–11 in) long, plus 22–25 cm (9–10 in) tail

Habitats: Forest

Diet: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Neacomys
(bristly mouse)

Brown mouse

Thomas, 1900

Eight species
Eastern Panama and South America Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (Dubost's bristly mouse) to 11 cm (4 in) long, plus 13 cm (5 in) tail (common bristly mouse)

Habitats: Forest

Diets: Seeds, insects, and fruit
Necromys
(bolo mouse)

Brown mouse

Ameghino, 1889

Nine species
South America Size range: 9 cm (4 in) long, plus 6 cm (2 in) tail (pleasant bolo mouse) to 15 cm (6 in) long, plus 11 cm (4 in) tail (northern grass mouse)

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland

Diets: Arthropods
Nectomys
(water rat)

Brown rat

Peters, 1861

Five species
South America and Trinidad and Tobago Size range: 11 cm (4 in) long, plus 12 cm (5 in) tail (South American water rat) to 29 cm (11 in) long, plus 25 cm (10 in) tail (common water rat)

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland

Diets: Vegetation, insects, tadpoles, and small fish
Neomicroxus
(small grass mouse)
Alvarado-Serrano & D'Elía, 2013

Two species
Northwestern South America Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (Bogotá grass mouse) to 11 cm (4 in) long, plus 10 cm (4 in) tail (Ecuadorian grass mouse)

Habitats: Grassland, forest, and shrubland

Diets: Vegetation, seeds, and insects
Neotomys

Brown rat

Thomas, 1894

One species
Western South America Size: 9–15 cm (4–6 in) long, plus 6–9 cm (2–4 in) tail

Habitats: Inland wetlands, shrubland, and grassland

Diet: Vegetation, seeds, and insects
Nephelomys
(misty rice rat)

Brown rat

Weksler, Percequillo, & Voss, 2006

Seven species
Central America and Northwestern South America Size range: 10 cm (4 in) long, plus 13 cm (5 in) tail (Tomes's rice rat) to 18 cm (7 in) long, plus 20 cm (8 in) tail (Ecuadorian rice rat)

Habitats: Grassland and forest

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Nesoryzomys
(Galápagos mouse)
Heller, 1904

Five species
Galápagos Islands Size range: 10 cm (4 in) long, plus 7 cm (3 in) tail (Fernandina Galápagos mouse) to 19 cm (7 in) long, plus 17 cm (7 in) tail (Santiago Galápagos mouse)

Habitats: Grassland, forest, and shrubland

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Neusticomys
(fish-eating rat)

Gray rat

Anthony, 1921

Six species
Northern South America Size range: 9–21 cm (4–8 in) long, plus 8–12 cm (3–5 in) tail (montane fish-eating rat)

Habitats: Savanna, forest, and inland wetlands

Diets: Aquatic invertebrates
Noronhomys † Olson & Carleton, 1999

One species
Fernando de Noronha islands northeast of Brazil
Map of range
Size: Unknown

Habitats: Shrubland

Diet: Vegetation, seeds, and insects
Notiomys Thomas, 1890

One species
Southern Argentina Size: 8–12 cm (3–5 in) long, plus 3–5 cm (1–2 in) tail

Habitats: Shrubland, grassland, and rocky areas

Diet: Insects and seeds
Oecomys
(arboreal rice rat)
Thomas, 1906

South America Size range: 7 cm (3 in) long, plus 8 cm (3 in) tail (red arboreal rice rat) to 17 cm (7 in) long, plus 19 cm (7 in) tail (foothill arboreal rice rat)

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Oligoryzomys
(pygmy rice rat)

Brown rat

Bangs, 1900

20 species
Central America and South America Size range: 7–12 cm (3–5 in) long, plus 10–15 cm (4–6 in) tail (black-footed pygmy rice rat)

Habitats: Inland wetlands, savanna, grassland, forest, desert, and shrubland

Diets: Seeds, fruit, and insects
Oreoryzomys

Brown and white rat

Weksler, Percequillo, & Voss, 2006

One species
Ecuador and northwestern Peru Size: 7–10 cm (3–4 in) long, plus 9–12 cm (4–5 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Oryzomys
(rice rat)

Gray rat

Baird, 1857

Six species
North America, Central America, South America, and Jamaica
Map of range
Size range: 9 cm (4 in) long, plus 10 cm (4 in) tail (Coues's rice rat) to 16 cm (6 in) long, plus 14 cm (6 in) tail (Gorgas's rice rat)

Habitats: Inland wetlands, grassland, forest, intertidal marine, and shrubland

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Oxymycterus
(hocicudo)

Brown rat

Waterhouse, 1837

Seventeen species
South America Size range: 8 cm (3 in) long, plus 10 cm (4 in) tail (Robert's hocicudo) to 20 cm (8 in) long, plus 16 cm (6 in) tail (angular hocicudo)

Habitats: Inland wetlands, savanna, grassland, forest, and shrubland

Diets: Insects, as well as other invertebrates and vegetation
Phaenomys

Brown rat

Thomas, 1917

One species
Southeastern Brazil Size: 14–17 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Phyllotis
(leaf-eared mouse)

Brown and white mouse

Waterhouse, 1837

Western and southern South America Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (gerbil leaf-eared mouse) to 16 cm (6 in) long, plus 15 cm (6 in) tail (Buenos Aires leaf-eared mouse)

Habitats: Rocky areas, inland wetlands, grassland, forest, desert, and shrubland

Diets: Seeds, vegetation, and lichen
Podoxymys Anthony, 1929

One species
Northern South America Size: 7–11 cm (3–4 in) long, plus 7–11 cm (3–4 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Pseudoryzomys Hershkovitz, 1962

One species
Central and eastern South America
Map of range
Size: 9–13 cm (4–5 in) long, plus 10–14 cm (4–6 in) tail

Habitats: Grassland, savanna, and shrubland

Diet: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Punomys
(puna mouse)
Osgood, 1943

Two species
Western South America Size range: 12–17 cm (5–7 in) long, plus 6–9 cm (2–4 in) tail (eastern puna mouse)

Habitats: Rocky areas, grassland, and shrubland

Diets: Twigs from herbs
Reithrodon
(bunny rat)

Brown rat

Waterhouse, 1837

Two species
Southeastern South America Size range: 12–15 cm (5–6 in) long, plus 7–10 cm (3–4 in) tail (bunny rat)

Habitats: Grassland and shrubland

Diets: Grass, rhizomes, and roots
Rhagomys
(arboreal mouse)
Thomas, 1917

Two species
Southern South America Size range: 8 cm (3 in) long, plus 8 cm (3 in) tail (Brazilian arboreal mouse) to 11 cm (4 in) long, plus 11 cm (4 in) tail (long-tongued arboreal mouse)

Habitats: Forest

Diets: Vegetation, seeds, and insects
Rheomys
(water mouse)

Brown and white rat head

Thomas, 1906

Four species
Southern Mexico and Central America Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Goldman's water mouse) to 15 cm (6 in) long, plus 18 cm (7 in) tail (Mexican water mouse)

Habitats: Forest and inland wetlands

Diets: Fish, snails, and aquatic insects
Rhipidomys
(climbing mouse)

Brown and white rat

Tschudi, 1845

22 species
Panama and South America Size range: 6 cm (2 in) long, plus 8 cm (3 in) tail (Wetzel's climbing mouse) to 21 cm (8 in) long, plus 20 cm (8 in) tail (Coues's climbing mouse)

Habitats: Rocky areas, savanna, grassland, and forest

Diets: Vegetation, seeds, and insects
Salinomys Braun & Mares, 1995

One species
Northern Argentina Size: 6–9 cm (2–4 in) long, plus 9–13 cm (4–5 in) tail

Habitats: Shrubland and inland wetlands

Diet: Seeds, arthropods, and vegetation
Scapteromys
(swamp rat)

Gray rat

Waterhouse, 1837

Two species
Southern South America Size range: 12–25 cm (5–10 in) long, plus 10–15 cm (4–6 in) tail (Argentine swamp rat)

Habitats: Grassland, forest, inland wetlands, and shrubland

Diets: Grass and seeds
Scolomys
(spiny mouse)
Anthony, 1924

Two species
Western South America Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (South American spiny mouse) to 12 cm (5 in) long, plus 8 cm (3 in) tail (Ucayali spiny mouse)

Habitats: Forest

Diets: Seeds and invertebrates
Sigmodon
(cotton rat)

Gray rat

Say & Ord, 1825

Fourteen species
North America, Central America, and Northern South America Size range: 10 cm (4 in) long, plus 7 cm (3 in) tail (Alston's cotton rat) to 24 cm (9 in) long, plus 16 cm (6 in) tail (Arizona cotton rat)

Habitats: Inland wetlands, grassland, forest, desert, and shrubland

Diets: Omnivorous, including vegetation, insects, invertebrates, and small animals
Sigmodontomys Allen, 1897

One species
Central America and northwestern South America Size: 12–18 cm (5–7 in) long, plus 14–19 cm (6–7 in) tail

Habitats: Inland wetlands and forest

Diet: Seeds and vegetation
Sooretamys Weksler, Percequillo, & Voss, 2006

One species
Southeastern South America Size: 12–21 cm (5–8 in) long, plus 16–24 cm (6–9 in) tail

Habitats: Forest and savanna

Diet: Vegetation, seeds, and insects
Tanyuromys Pine, Timm, & Weksler, 2012

One species
Southern Central America and northwestern South America Size: 11–16 cm (4–6 in) long, plus 17–23 cm (7–9 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Tapecomys

Brown rat

Anderson & Yates, 2000

One species
Southern Bolivia and northern Argentina Size: 12–16 cm (5–6 in) long, plus 14–16 cm (6 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Thalpomys
(cerrado mouse)

Brown mouse

Thomas, 1916

Two species
Brazil Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (hairy-eared cerrado mouse) to 11 cm (4 in) long, plus 7 cm (3 in) tail (Cerrado mouse)

Habitats: Savanna, grassland, and shrubland

Diets: Vegetation, seeds, and insects
Thaptomys

Gray mouse

Thomas, 1916

One species
Southeastern South America Size: 8–10 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail

Habitats: Grassland, shrubland, and forest

Diet: Plants and invertebrates
Thomasomys
(Oldfield mouse)

Brown mouse

Coues, 1884

42 species
Northwestern South America Size range: 8 cm (3 in) long, plus 14 cm (6 in) tail (Austral Oldfield mouse) to 24 cm (9 in) long, plus 33 cm (13 in) tail (Apeco Oldfield mouse)

Habitats: Rocky areas, inland wetlands, grassland, forest, and shrubland

Diets: Vegetation, seeds, and insects
Transandinomys
(transandean rice rat)
Weksler, Percequillo, & Voss, 2006

Two species
Central America and northwestern South America Size range: 10 cm (4 in) long, plus 9 cm (4 in) tail (long-whiskered rice rat) to 16 cm (6 in) long, plus 16 cm (6 in) tail (Talamancan rice rat)

Habitats: Forest and inland wetlands

Diets: Grass, sedges, seeds, fruit, insects, crustaceans, and small fish
Wiedomys
(red-nosed mouse)

Brown and white mouse

Hershkovitz, 1959

Two species
Eastern Brazil Size range: 10 cm (4 in) long, plus 14 cm (6 in) tail (Cerrado red-nosed mouse) to 12 cm (5 in) long, plus 19 cm (7 in) tail (red-nosed mouse)

Habitats: Savanna, grassland, forest, and shrubland

Diets: Seeds and insects
Wilfredomys

Brown mouse

Avila-Pires, 1960

One species
Southeastern South America Size: 11–13 cm (4–5 in) long, plus 16–21 cm (6–8 in) tail

Habitats: Forest

Diet: Vegetation, seeds, and insects
Zygodontomys
(cane mouse)
Allen, 1897

Two species
Southern Central America and northern South America Size range: 13 cm (5 in) long, plus 11 cm (4 in) tail (brown cane mouse) to 17 cm (7 in) long, plus 12 cm (5 in) tail (short-tailed cane mouse)

Habitats: Savanna, grassland, forest, and inland wetlands

Diets: Seeds, grass, and fruit
Subfamily Tylomyinae – Reig, 1984 – four genera
Name Authority and species Range Size and ecology
Nyctomys

Brown rat

Saussure, 1860

One species
Southern Mexico, Central America
Map of range
Size: 12–14 cm (5–6 in) long, plus 14–16 cm (6 in) tail

Habitats: Forest

Diet: Seeds, fruit, and other vegetation
Otonyctomys

Brown rat

Anthony, 1932

One species
Eastern Mexico and northern Central America
Map of range
Size: 9–12 cm (4–5 in) long, plus 6–13 cm (2–5 in) tail

Habitats: Forest

Diet: Vegetation
Ototylomys

Gray rat

Merriam, 1901

One species
Mexico and Central America Size: 12–17 cm (5–7 in) long, plus 10–18 cm (4–7 in) tail

Habitats: Forest

Diet: Fruit and leaves
Tylomys
(climbing rat)

Gray rat

Peters, 1866

Seven species
Mexico, Central America, and northwestern South America Size range: 15–29 cm (6–11 in) long, plus 15–32 cm (6–13 in) tail (Watson's climbing rat)

Habitats: Forest

Diets: Seeds, fruit, and other vegetation
Family Muridae

Members of the Muridae family are called murids and include gerbils and many types of mice and rats. They eat a variety of vegetation and invertebrates, though some also eat small vertebrates. Muridae comprises 801 extant species, the most of any mammal family, divided into 156 genera. These genera are divided into five subfamilies: Deomyinae, containing 55 species of spiny mice and brush-furred rats; Gerbillinae, containing 102 species of gerbils, jirds, and sand rats; Leimacomyinae, containing a single rat species; Lophiomyinae, containing a single mouse species; and Murinae, containing 642 species of Old World mice and rats. Additionally, thirteen species in Murinae have been driven extinct since 1500.

Subfamily Deomyinae – Thomas, 1888 – four genera
Name Authority and species Range Size and ecology
Acomys
(spiny mouse)

Light brown mouse

I. Geoffroy, 1838

22 species
Africa, southern Europe, and southwestern Asia Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (Cape spiny mouse) to 14 cm (6 in) long, plus 12 cm (5 in) tail (Cyprus spiny mouse)

Habitats: Savanna, forest, shrubland, grassland, desert, and rocky areas

Diets: Omnivorous, but primarily grain, grass, and other plant material
Deomys Thomas, 1888

One species
Central Africa Size: 10–15 cm (4–6 in) long, plus 15–20 cm (6–8 in) tail

Habitats: Inland wetlands and forest

Diet: Insects and other invertebrates, as well as vegetable matter
Lophuromys
(brush-furred rat)

Brown mouse

Peters, 1874

Western, central, and eastern Africa Size range: 8 cm (3 in) long, plus 5 cm (2 in) tail (Dudu's brush-furred rat) to 18 cm (7 in) long, plus 10 cm (4 in) tail (black-clawed brush-furred rat)

Habitats: Rocky areas, grassland, forest, and shrubland

Diets: Insects, as well as other invertebrates, frogs, other small vertebrates, and vegetation
Uranomys

Brown mouse

Dollman, 1909

One species
Western, central, and eastern Africa Size: 9–12 cm (4–5 in) long, plus 6–8 cm (2–3 in) tail

Habitats: Savanna and shrubland

Diet: Insects
Subfamily Gerbillinae – Gray, 1825 – fourteen genera
Name Authority and species Range Size and ecology
Ammodillus Thomas, 1904

One species
Ethiopia and Somalia Size: 8–11 cm (3–4 in) long, plus 12–15 cm (5–6 in) tail

Habitats: Grassland

Diet: Plant material and invertebrates
Brachiones

Drawing of brown gerbils

Thomas, 1925

One species
Northern China Size: 6–11 cm (2–4 in) long, plus 5–8 cm (2–3 in) tail

Habitats: Desert

Diet: Plant material and invertebrates
Desmodilliscus Wettstein, 1916

One species
Western and northern Africa
Map of range
Size: 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail

Habitats: Savanna

Diet: Grain
Desmodillus

Brown gerbil

Thomas & Schwann, 1904

One species
Southern Africa Size: 8–13 cm (3–5 in) long, plus 7–10 cm (3–4 in) tail

Habitats: Desert

Diet: Seeds, grain, and insects
Gerbilliscus
(large naked-soled gerbil)

Brown gerbil

Thomas, 1897

Sixteen species
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 12 cm (5 in) tail (bushveld gerbil) to 20 cm (8 in) long, plus 21 cm (8 in) tail (black-tailed gerbil)

Habitats: Desert, forest, savanna, grassland, shrubland, and rocky areas

Diets: Roots, bulbs, seeds, vegetation, and insects; Seeds
Gerbillus
(northern pygmy gerbil)

Brown and white gerbil

Desmarest, 1804

49 species
Northern, central, and eastern Africa and western and southern Asia Size range: 5 cm (2 in) long, plus 8 cm (3 in) tail (least gerbil) to 22 cm (9 in) long, plus 11 cm (4 in) tail (large Aden gerbil)

Habitats: Desert, savanna, forest, grassland, shrubland, coastal marine, and rocky areas

Diets: Seeds, roots, nuts, grass, and insects
Meriones
(jird)

Brown and white gerbil

Illiger, 1811

Seventeen species
Northern Africa and western, central, and southern Asia Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Mongolian gerbil) to 20 cm (8 in) long, plus 18 cm (7 in) tail (Moroccan jird)

Habitats: Desert, savanna, grassland, inland wetlands, shrubland, and rocky areas

Diets: Vegetation, roots, bulbs, seeds, grain, fruit, and insects
Microdillus Thomas, 1910

One species
Somalia Size: 6–8 cm (2–3 in) long, plus 5–7 cm (2–3 in) tail

Habitats: Grassland and shrubland

Diet: Seeds, roots, nuts, grass, and insects
Pachyuromys

Brown gerbils

Lataste, 1880

One species
Northern Africa Size: 9–12 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail

Habitats: Shrubland and desert

Diet: Insects
Psammomys
(sand rat)

Brown gerbil

Cretzschmar, 1828

Two species
Northern Africa and Middle East Size range: 11 cm (4 in) long, plus 8 cm (3 in) tail (thin sand rat) to 19 cm (7 in) long, plus 14 cm (6 in) tail (fat sand rat)

Habitats: Inland wetlands, desert, and shrubland

Diets: Succulents and other plants
Rhombomys

Brown gerbil

Wagner, 1841

One species
Central Asia Size: 15–19 cm (6–7 in) long, plus 13–16 cm (5–6 in) tail

Habitats: Desert, grassland, and shrubland

Diet: Variety of desert plants
Sekeetamys

Brown gerbil

Ellerman, 1947

One species
Egypt and Middle East Size: 9–13 cm (4–5 in) long, plus 11–17 cm (4–7 in) tail

Habitats: Rocky areas and desert

Diet: Plants and insects
Tatera

Brown and white gerbil

Lataste, 1882

One species
Southern and western Asia Size: 14–21 cm (6–8 in) long, plus 15–20 cm (6–8 in) tail

Habitats: Forest, desert, grassland, and shrubland

Diet: Roots, bulbs, seeds, vegetation, insects, eggs, and birds
Taterillus
(tateril)

Brown and white gerbil

Thomas, 1910

Nine species
Eastern, central, and western Africa Size range: 9 cm (4 in) long, plus 13 cm (5 in) tail (gracile tateril) to 15 cm (6 in) long, plus 19 cm (7 in) tail (Congo gerbil)

Habitats: Grassland, forest, savanna, and shrubland

Diets: Grain and insects
Subfamily Leimacomyinae – Musser & Carleton, 2005 – one genus
Name Authority and species Range Size and ecology
Leimacomys Matschie, 1893

One species
Togo
Map of range
Size: About 12 cm (5 in) long, plus about 4 cm (2 in) tail

Habitats: Unknown

Diet: Vegetation and insects
Subfamily Lophiomyinae – Milne-Edwards, 1867 – one genus
Name Authority and species Range Size and ecology
Lophiomys

Light brown mouse

Milne-Edwards, 1867

One species
Eastern Africa Size: 25–30 cm (10–12 in) long, plus 14–20 cm (6–8 in) tail

Habitats: Forest, savanna, and shrubland

Diet: Leaves and shoots
Subfamily Murinae – Illiger, 1811 – 136 genera
Name Authority and species Range Size and ecology
Abditomys Musser, 1982

One species
Philippines
Map of range
Size: 23–27 cm (9–11 in) long, plus 22–24 cm (9 in) tail

Habitats: Forest

Diet: Vegetation
Abeomelomys Menzies, 1990

One species
Papua New Guinea Size: 11–14 cm (4–6 in) long, plus 14–20 cm (6–8 in) tail

Habitats: Grassland and forest

Diet: Plant material and invertebrates
Aethomys
(rock rat)

Brown rat

Thomas, 1915

Nine species
Central, eastern, and southern Africa Size range: 12 cm (5 in) long, plus 12 cm (5 in) tail (Hinde's rock rat) to 20 cm (8 in) long, plus 18 cm (7 in) tail (Bocage's rock rat)

Habitats: Savanna, grassland, forest, rocky areas, and shrubland

Diets: Grain, seeds, roots, nuts, and fruit
Anisomys Thomas, 1904

One species
New Guinea Size: 21–28 cm (8–11 in) long, plus 27–36 cm (11–14 in) tail

Habitats: Forest

Diet: Nuts and other vegetation
Anonymomys Musser, 1981

One species
Philippines
Map of range
Size: 14–15 cm (6 in) long, plus 19–22 cm (7–9 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Apodemus
(field mouse)

Brown and white mouse

Kaup, 1829

20 species
Europe, Asia, and northern Africa Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (small Japanese field mouse) to 15 cm (6 in) long, plus 15 cm (6 in) tail (western broad-toothed field mouse)

Habitats: Coastal marine, grassland, inland wetlands, forest, rocky areas, and shrubland

Diets: Roots, grain, seeds, berries, nuts, and insects
Apomys
(earthworm mouse)

Stamp with brown mouse

Mearns, 1905

Nineteen species
Philippines Size range: 7 cm (3 in) long, plus 9 cm (4 in) tail (least forest mouse) to 18 cm (7 in) long, plus 16 cm (6 in) tail (Sierra Madre forest mouse)

Habitats: Forest, shrubland, and grassland

Diets: Omnivorous
Archboldomys
(shrew-mouse)
Musser, 1982

Two species
Philippines Size range: 10–12 cm (4–5 in) long, plus 9–11 cm (4 in) tail (large Cordillera shrew-mouse)

Habitats: Forest

Diets: Worms and soft-bodied invertebrates
Arvicanthis
(unstriped grass rat)

Brown rat

Lesson, 1842

Seven species
Africa Size range: 11 cm (4 in) long, plus 12 cm (5 in) tail (Guinean grass rat) to 19 cm (7 in) long, plus 16 cm (6 in) tail (African grass rat)

Habitats: Savanna, shrubland, and grassland

Diets: Seeds, leaves, grass, and grain
Baiyankamys
(mountain water rat)
Hinton, 1943

Two species
New Guinea
Map of range
Size range: 13–16 cm (5–6 in) long, plus 15–19 cm (6–7 in) tail (mountain water rat)

Habitats: Forest, inland wetlands, grassland, and shrubland

Diets: Fish and aquatic insects, as well as spiders, crustaceans, mussels, frogs, turtles, birds, and bats
Bandicota
(bandicoot rat)

Brown rat

Gray, 1873

Three species
Southern and southeastern Asia Size range: 14 cm (6 in) long, plus 7 cm (3 in) tail (Savile's bandicoot rat) to 35 cm (14 in) long, plus 28 cm (11 in) tail (greater bandicoot rat)

Habitats: Forest, inland wetlands, shrubland, and grassland

Diets: Omnivorous
Batomys
(hairy-tailed rat)

Gray rat

Thomas, 1895

Six species
Philippines
Map of range
Size range: 13 cm (5 in) long, plus 13 cm (5 in) tail (Mindanao hairy-tailed rat) to 21 cm (8 in) long, plus 18 cm (7 in) tail (Mount Isarog hairy-tailed rat)

Habitats: Forest

Diets: Leaves, seeds, and fruit
Berylmys
(white-toothed rat)

Drawing of brown rat

Ellerman, 1947

Five species
Southeastern Asia and eastern China Size range: 13 cm (5 in) long, plus 14 cm (6 in) tail (Manipur white-toothed rat) to 30 cm (12 in) long, plus 31 cm (12 in) tail (Bower's white-toothed rat)

Habitats: Forest, grassland, and shrubland

Diets: Leaves, grass, seeds, fruit, insects, molluscs, and worms
Brassomys Musser & Lunde, 2009

One species
Western New Guinea
Map of range
Size: 11–13 cm (4–5 in) long, plus 14–17 cm (6–7 in) tail

Habitats: Grassland and shrubland

Diet: Leaves and other vegetation
Bullimus
(large forest rat)

Gray rat

Mearns, 1905

Three species
Philippines Size range: 22 cm (9 in) long, plus 14 cm (6 in) tail (Camiguin forest rat) to 27 cm (11 in) long, plus 20 cm (8 in) tail (Bagobo rat)

Habitats: Forest and shrubland

Diets: Plant material and invertebrates
Bunomys
(hill rat)

Dead brown rat

Thomas, 1910

Seven species
Indonesia Size range: 9 cm (4 in) long, plus 9 cm (4 in) tail (yellow-haired hill rat) to 24 cm (9 in) long, plus 19 cm (7 in) tail (inland hill rat)

Habitats: Forest and shrubland

Diets: Fruit, insects, snails, and worms
Carpomys
(Luzon tree rat)

Drawing of brown and white rat

Thomas, 1895

Two species
Philippines Size range: 16 cm (6 in) long, plus 15 cm (6 in) tail (white-bellied Luzon tree rat) to 19 cm (7 in) long, plus 19 cm (7 in) tail (short-footed Luzon tree rat)

Habitats: Forest

Diets: Plant material and invertebrates
Chiromyscus Thomas, 1925

One species
Southeastern Asia
Map of range
Size: 13–16 cm (5–6 in) long, plus 20–23 cm (8–9 in) tail

Habitats: Shrubland and forest

Diet: Plant material and invertebrates
Chiropodomys
(pencil-tailed tree mouse)
Peters, 1869

Six species
Southeastern Asia Size range: 6 cm (2 in) long, plus 8 cm (3 in) tail (gray-bellied pencil-tailed tree mouse) to 13 cm (5 in) long, plus 18 cm (7 in) tail (Palawan pencil-tailed tree mouse)

Habitats: Forest

Diets: Vegetation
Chiruromys
(tree mouse)
Thomas, 1888

Three species
Papua New Guinea Size range: 8 cm (3 in) long, plus 12 cm (5 in) tail (lesser tree mouse) to 17 cm (7 in) long, plus 23 cm (9 in) tail (greater tree mouse)

Habitats: Forest

Diets: Leaves and grass and bamboo shoots
Chrotomys
(striped shrew-rat)

Drawing of brown, yellow, and white rat

Thomas, 1895

Five species
Philippines Size range: 13 cm (5 in) long, plus 9 cm (4 in) tail (blazed Luzon shrew-rat) to 19 cm (7 in) long, plus 11 cm (4 in) tail (Isarog striped shrew-rat)

Habitats: Forest

Diets: Sweet potatoes, grass, and worms
Coccymys
(brush mouse)
Menzies, 1990

Three species
Papua New Guinea and New Guinea Size range: 8 cm (3 in) long, plus 13 cm (5 in) tail (Central Cordillera brush mouse) to 12 cm (5 in) long, plus 18 cm (7 in) tail (Rümmler's brush mouse)

Habitats: Forest and grassland

Diets: Leaves and other vegetation
Colomys Thomas & Wroughton, 1907

One species
Central Africa Size: 10–16 cm (4–6 in) long, plus 14–19 cm (6–7 in) tail

Habitats: Savanna, forest, grassland, and inland wetlands

Diet: Worms, slugs, crustaceans, and aquatic insects, as well as small vertebrates and some vegetation
Conilurus
(rabbit rat)

Drawing of brown and white rat

Ogilby, 1838

Three species
Southeastern Australia, Northern Australia and Papua New Guinea, and Northeastern Australia Size range: 15–20 cm (6–8 in) long, plus 17–21 cm (7–8 in) tail (brush-tailed rabbit rat)

Habitats: Savanna and forest

Diets: Plant material and invertebrates
Crateromys
(bushy-tailed cloud rat)

Brown rat

Thomas, 1895

Four species
Philippines Size range: 25 cm (10 in) long, plus 21 cm (8 in) tail (Ilin Island cloudrunner) to 37 cm (15 in) long, plus 39 cm (15 in) tail (giant bushy-tailed cloud rat)

Habitats: Forest

Diets: Fruit and pine tree sprouts, buds, and bark
Cremnomys
(Cutch rat)

Brown rat

Wroughton, 1912

Two species
India and Southern India Size range: 10 cm (4 in) long, plus 12 cm (5 in) tail (Cutch rat) to 15 cm (6 in) long, plus 20 cm (8 in) tail (Elvira rat)

Habitats: Grassland, desert, forest, rocky areas, and shrubland

Diets: Plant material and invertebrates
Crossomys Thomas, 1907

One species
New Guinea
Map of range
Size: 17–23 cm (7–9 in) long, plus 21–27 cm (8–11 in) tail

Habitats: Forest and inland wetlands

Diet: Tadpoles, insects, mollusks, and small aquatic vertebrates
Crunomys
(shrew-rat)

Drawing of brown and white rat

Thomas, 1897

Four species
Philippines and Indonesia Size range: 9–14 cm (4–6 in) long, plus 6–10 cm (2–4 in) tail (Mindanao shrew-rat)

Habitats: Forest

Diets: Plant material and invertebrates
Dacnomys Thomas, 1916

One species
Southeastern Asia
Map of range
Size: 21–27 cm (8–11 in) long, plus 29–33 cm (11–13 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Dasymys
(marsh rat)
Peters, 1875

Nine species
Sub-Saharan Africa
Map of range
Size range: 11 cm (4 in) long, plus 10 cm (4 in) tail (Glover Allen's dasymys) to 20 cm (8 in) long, plus 16 cm (6 in) tail (African marsh rat)

Habitats: Savanna, grassland, inland wetlands, forest, and shrubland

Diets: Aquatic plants, as well as insects
Dephomys
(Ivory Coast rat)
Thomas, 1926

Two species
Western Africa Size range: 11 cm (4 in) long, plus 18 cm (7 in) tail (Defua rat) to 15 cm (6 in) long, plus 20 cm (8 in) tail (Ivory Coast rat)

Habitats: Forest

Diets: Fruit and insects
Desmomys
(scrub rat)
Thomas, 1910

Two species
Ethiopia Size range: 11 cm (4 in) long, plus 14 cm (6 in) tail (Yalden's rat) to 15 cm (6 in) long, plus 14 cm (6 in) tail (Harrington's rat)

Habitats: Forest and shrubland

Diets: Plant material and invertebrates
Diomys Thomas, 1917

One species
Scattered southern Asia
Map of range
Size: 10–14 cm (4–6 in) long, plus 10–13 cm (4–5 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Diplothrix

Brown rat

Thomas, 1916

One species
Southern islands of Japan Size: About 23 cm (9 in)

Habitats: Forest

Diet: Plant material and invertebrates
Echiothrix
(echiothrix)

Dead brown rat

Gray, 1867

Two species
Indonesia Size range: 18 cm (7 in) long, plus 23 cm (9 in) tail (central Sulawesi echiothrix) to 23 cm (9 in) long, plus 27 cm (11 in) tail (northern Sulawesi echiothrix)

Habitats: Forest

Diets: Worms
Eropeplus Miller & Hollister, 1921

One species
Indonesia Size: 23–25 cm (9–10 in) long, plus 27–30 cm (11–12 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Frateromys Sody, 1941

One species
Indonesia Size: 15–19 cm (6–7 in) long, plus 15–20 cm (6–8 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Golunda

Brown rat

Gray, 1837

One species
Southern Asia Size: 9–17 cm (4–7 in) long, plus 7–14 cm (3–6 in) tail

Habitats: Shrubland, forest, and grassland

Diet: Roots and grass, as well as other vegetation
Gracilimus

Brown rat

Rowe, Achmadi, & Esselstyn, 2016

One species
Indonesia Size: 10–13 cm (4–5 in) long, plus 16–17 cm (6–7 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Grammomys
(thicket rat)

Gray rat

Thomas, 1915

Eleven species
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 14 cm (6 in) tail (gray-headed thicket rat) to 15 cm (6 in) long, plus 18 cm (7 in) tail (eastern rainforest grammomys)

Habitats: Savanna, grassland, inland wetlands, forest, and shrubland

Diets: Stems, fruit, nuts, flowers, and other vegetation, as well as insects
Hadromys
(bush rat)

Drawing of brown and white rat

Thomas, 1911

Two species
Eastern India and Southern China Size range: 9–14 cm (4–6 in) long, plus 11–14 cm (4–6 in) tail (multiple)

Habitats: Forest

Diets: Grass
Haeromys
(ranee mouse)
Thomas, 1911

Three species
Southeastern Asia Size range: 7 cm (3 in) long, plus 12 cm (5 in) tail (lesser ranee mouse) to 8 cm (3 in) long, plus 14 cm (6 in) tail (Minahassa ranee mouse)

Habitats: Forest

Diets: Seeds
Halmaheramys
(Halmahera spiny rat)
Fabre, Pagès, Musser, Fitriana, Semiadi, & Helgen, 2013

Two species
Indonesia Size range: 14 cm (6 in) long, plus 11 cm (4 in) tail (spiny Boki Mekot rat) to 24 cm (9 in) long, plus 21 cm (8 in) tail (Wallace's large spiny rat)

Habitats: Forest

Diets: Plant material and invertebrates
Hapalomys
(marmoset rat)

Brown rat

Blyth, 1859

Three species
Southeastern Asia Size range: 10 cm (4 in) long, plus 13 cm (5 in) tail (Delacour's marmoset rat) to 17 cm (7 in) long, plus 20 cm (8 in) tail (marmoset rat)

Habitats: Forest

Diets: Bamboo shoots, flowers, and fruit
Heimyscus Misonne, 1969

One species
West-central Africa Size: 8–10 cm (3–4 in) long, plus 8–12 cm (3–5 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Hybomys
(striped mouse)
Thomas, 1910

Six species
Western Uganda and eastern Democratic Republic of the Congo, Central Africa, Western Africa, and Cameroon Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (Eisentraut's striped mouse) to 16 cm (6 in) long, plus 14 cm (6 in) tail (Father Basilio's striped mouse)

Habitats: Forest

Diets: Fruit, roots, and insects
Hydromys
(water rat)

Brown rat

Geoffroy, 1804

Four species
Australia and southeastern Asia Size range: 12 cm (5 in) long, plus 10 cm (4 in) tail (western water rat) to 39 cm (15 in) long, plus 32 cm (13 in) tail (rakali)

Habitats: Coastal marine, forest, inland wetlands, and neritic marine

Diets: Fish and aquatic insects, as well as spiders, crustaceans, mussels, frogs, turtles, birds, and bats
Hylomyscus
(wood mouse)
Thomas, 1926

Sixteen species
Eastern, central, and western Africa Size range: 5 cm (2 in) long, plus 5 cm (2 in) tail (Dahomey Gap wood mouse) to 12 cm (5 in) long, plus 15 cm (6 in) tail (Baer's wood mouse)

Habitats: Forest

Diets: Fruit and other vegetation, as well as animals
Hyomys
(white-eared giant rat)
Thomas, 1904

Two species
New Guinea Size range: 29 cm (11 in) long, plus 24 cm (9 in) tail (western white-eared giant rat) to 39 cm (15 in) long, plus 38 cm (15 in) tail (eastern white-eared giant rat)

Habitats: Forest

Diets: Shoots as well as other vegetation
Hyorhinomys

Brown and white rat

Esselstyn, Achmadi, Handika, & Rowe, 2015

One species
Indonesia Size: 11–13 cm (4–5 in) long, plus 8–10 cm (3–4 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Kadarsanomys Musser, 1981

One species
Indonesia
Map of range
Size: 16–21 cm (6–8 in) long, plus 25–30 cm (10–12 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Komodomys

Dead brown rat

Musser & Boeadi, 1980

One species
Indonesia Size: 12–18 cm (5–7 in) long, plus 12–17 cm (5–7 in) tail

Habitats: Shrubland and forest

Diet: Plant material and invertebrates
Lamottemys Petter, 1986

One species
Cameroon Size: 13–15 cm (5–6 in) long, plus 12–14 cm (5–6 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Leggadina
(short-tailed mouse)

Dead brown mouse

Thomas, 1910

Two species
Northern and central Australia Size range: 5 cm (2 in) long, plus 4 cm (2 in) tail (Lakeland Downs mouse) to 10 cm (4 in) long, plus 7 cm (3 in) tail (Forrest's mouse)

Habitats: Desert, savanna, grassland, and shrubland

Diets: Seeds and vegetation
Lemniscomys
(striped grass mouse)

Brown and white mouse

Trouessart, 1881

Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 7 cm (3 in) tail (buffoon striped grass mouse) to 17 cm (7 in) long, plus 16 cm (6 in) tail (single-striped grass mouse)

Habitats: Savanna, forest, grassland, and shrubland

Diets: Grass, seeds, grains, and insects
Lenomys Thomas, 1898

One species
Indonesia Size: 23–30 cm (9–12 in) long, plus 24–30 cm (9–12 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Lenothrix

Dead brown rat

Miller, 1903

One species
Malaysia and Brunei Size: 14–21 cm (6–8 in) long, plus 19–29 cm (7–11 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Leopoldamys
(long-tailed giant rat)

Brown and white rat

Ellerman, 1947

Seven species
Southeastern Asia Size range: 17 cm (7 in) long, plus 22 cm (9 in) tail (Mentawai long-tailed giant rat) to 29 cm (11 in) long, plus 31 cm (12 in) tail (Edwards's long-tailed giant rat)

Habitats: Forest

Diets: Insects, other invertebrates, and a wide variety of vegetation
Leporillus
(stick-nest rat)

Brown rat

Thomas, 1906

Two species
Southern and western Australia Size range: 17–26 cm (7–10 in) long, plus 14–18 cm (6–7 in) tail (greater stick-nest rat)

Habitats: Savanna, grassland, desert, caves, rocky areas, and shrubland

Diets: Succulents and other vegetation
Leptomys
(narrow water rat)
Thomas, 1897

Five species
New Guinea Size range: 11 cm (4 in) long, plus 13 cm (5 in) tail (small water rat) to 20 cm (8 in) long, plus 17 cm (7 in) tail (long-footed water rat)

Habitats: Forest

Diets: Insects and small animals
Limnomys
(mountain rat)

Dead brown rat

Mearns, 1905

Two species
Philippines Size range: 11–15 cm (4–6 in) long, plus 14–19 cm (6–7 in) tail (multiple)

Habitats: Forest

Diets: Plant material and invertebrates
Lorentzimys Jentink, 1911

One species
New Guinea Size: 5–9 cm (2–4 in) long, plus 11–13 cm (4–5 in) tail

Habitats: Forest

Diet: Insects, vegetation, and fungi
Macruromys
(small-toothed rat)
Stein, 1933

Two species
Western New Guinea and New Guinea Size range: 15 cm (6 in) long, plus 21 cm (8 in) tail (lesser small-toothed rat) to 26 cm (10 in) long, plus 34 cm (13 in) tail (eastern small-toothed rat)

Habitats: Forest

Diets: Vegetation
Madromys

Brown and white rat

Sody, 1941

One species
Southern Asia Size: 15–19 cm (6–7 in) long, plus 18–21 cm (7–8 in) tail

Habitats: Caves, forest, and shrubland

Diet: Plant material and invertebrates
Malacomys
(swamp rat)

Brown and white rat

A. Milne-Edwards, 1877

Three species
Central and western Africa Size range: 12 cm (5 in) long, plus 15 cm (6 in) tail (Edward's swamp rat) to 18 cm (7 in) long, plus 20 cm (8 in) tail (big-eared swamp rat)

Habitats: Forest, inland wetlands, and shrubland

Diets: Fruit, seeds, nuts, roots, insects, slugs, snails, and crabs
Mallomys
(woolly rat)

Gray rat

Thomas, 1898

Four species
New Guinea Size range: 34 cm (13 in) long, plus 33 cm (13 in) tail (De Vis's woolly rat) to 47 cm (19 in) long, plus 37 cm (15 in) tail (alpine woolly rat)

Habitats: Grassland, forest, caves, rocky areas, and shrubland

Diets: Shoots as well as other vegetation
Mammelomys
(large mosaic-tailed rat)
Menzies, 1996

Two species
New Guinea Size range: 13 cm (5 in) long, plus 10 cm (4 in) tail (large-scaled mosaic-tailed rat) to 22 cm (9 in) long, plus 15 cm (6 in) tail (large mosaic-tailed rat)

Habitats: Forest

Diets: Plant material and invertebrates
Margaretamys
(margareta rat)

Dead brown rat

Musser, 1981

Four species
Indonesia Size range: 9 cm (4 in) long, plus 15 cm (6 in) tail (little margareta rat) to 20 cm (8 in) long, plus 29 cm (11 in) tail (elegant margareta rat)

Habitats: Forest

Diets: Plant material and invertebrates
Mastacomys

Gray mouse

Thomas, 1882

One species
Southeastern Australia Size: 14–20 cm (6–8 in) long, plus 10–14 cm (4–6 in) tail

Habitats: Forest, shrubland, grassland, and inland wetlands

Diet: Plant material and invertebrates
Mastomys
(multimammate mouse)

Brown and white mouse

Thomas, 1915

Eight species
Morocco and Sub-Saharan Africa Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (dwarf multimammate mouse) to 18 cm (7 in) long, plus 16 cm (6 in) tail (Guinea multimammate mouse)

Habitats: Savanna, grassland, inland wetlands, forest, and shrubland

Diets: Grass, seeds, and insects
Maxomys
(spiny rat)

Brown and white rat

Sody, 1936

Eighteen species
Southeastern Asia Size range: 9 cm (4 in) long, plus 10 cm (4 in) tail (Tajuddin's spiny rat) to 23 cm (9 in) long, plus 21 cm (8 in) tail (rajah spiny rat)

Habitats: Forest and shrubland

Diets: Roots, fruit, and other vegetation, as well as invertebrates and small vertebrates
Melasmothrix Miller & Hollister, 1921

One species
Indonesia Size: 11–13 cm (4–5 in) long, plus 8–10 cm (3–4 in) tail

Habitats: Forest

Diet: Worms and insect larvae
Melomys
(mosaic-tailed rat)
Thomas, 1922

Southeastern Asia and Australia Size range: 9–20 cm (4–8 in) long, plus 10–18 cm (4–7 in) tail (fawn-footed mosaic-tailed rat)

Habitats: Savanna, grassland, forest, inland wetlands, and shrubland

Diets: Fruit, berries, and other vegetation
Mesembriomys
(tree-rat)

Drawing of brown and white rat

Palmer, 1906

Two species
Northern Australia Size range: 18 cm (7 in) long, plus 29 cm (11 in) tail (golden-backed tree-rat) to 30 cm (12 in) long, plus 41 cm (16 in) tail (black-footed tree-rat)

Habitats: Savanna and forest

Diets: Seeds, nuts, and insects
Micaelamys
(rock mouse)

Brown rat

Ellerman, 1941

Two species
Southern Africa Size range: 8–15 cm (3–6 in) long, plus 10–20 cm (4–8 in) tail (Namaqua rock rat)

Habitats: Savanna, grassland, desert, forest, rocky areas, and shrubland

Diets: Grain, seeds, roots, nuts, and fruit
Microhydromys
(groove-toothed shrew mouse)
Tate & Archbold, 1941

Two species
New Guinea Size range: 7–9 cm (3–4 in) long, plus 8–10 cm (3–4 in) tail (northern groove-toothed shrew mouse)

Habitats: Savanna and forest

Diets: Insects
Micromys

Brown mouse

Dehne, 1841

One species
Europe and Asia
Map of range
Size: 4–8 cm (2–3 in) long, plus 3–8 cm (1–3 in) tail

Habitats: Forest and inland wetlands

Diet: Seeds, vegetation, and insects, as well as bird eggs
Millardia
(soft-furred rat)

Brown rat

Thomas, 1911

Four species
Southern Asia Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (sand-colored soft-furred rat) to 20 cm (8 in) long, plus 19 cm (7 in) tail (Kondana rat)

Habitats: Grassland, desert, forest, rocky areas, and shrubland

Diets: Grain, seeds, and swamp vegetation
Mirzamys
(moss rat)
Helgen & Helgen, 2009

Two species
New Guinea Size range: 9 cm (4 in) long, plus 10 cm (4 in) tail (Mirza's western moss rat) to 12 cm (5 in) long, plus 10 cm (4 in) tail (Mirza's eastern moss rat)

Habitats: Forest and grassland

Diets: Plant material and invertebrates
Muriculus Rüppell, 1842

One species
Ethiopia Size: 7–8 cm (3 in) long, plus 4–6 cm (2 in) tail

Habitats: Grassland

Diet: Plant material and invertebrates
Mus
(mouse)

Brown and white mouse

Linnaeus, 1758

39 species
Africa, Europe, and Asia Size range: 4 cm (2 in) long, plus 2 cm (0.8 in) tail (African pygmy mouse) to 13 cm (5 in) long, plus 9 cm (4 in) tail (stone-loving mouse)

Habitats: Grassland

Diets: Seeds, roots, leaves, stems, and other vegetation, as well as insects and some meat
Musseromys
(tree-mouse)
Heaney, Balete, Rickart, Veluz, & Jansa, 2009

Four species
Philippines Size range: 7–9 cm (3–4 in) long, plus 8 cm (3 in) tail (Mount Pulag tree-mouse)

Habitats: Forest

Diets: Plant material and invertebrates
Mylomys
(groove-toothed rat)
Thomas, 1906

Two species
Northern Sub-Saharan Africa Size range: 12 cm (5 in) long, plus 10 cm (4 in) tail (African groove-toothed rat) to 21 cm (8 in) long, plus 18 cm (7 in) tail (Ethiopian mylomys)

Habitats: Savanna and forest

Diets: Grass and leaves
Myomyscus
(white-footed rat)

Brown mouse

Shortridge, 1942

Four species
Sub-Saharan Africa, southern Saudi Arabia and Yemen Size range: 7 cm (3 in) long, plus 15 cm (6 in) tail (Yemeni mouse) to 13 cm (5 in) long, plus 18 cm (7 in) tail (Brockman's rock mouse)

Habitats: Savanna, rocky areas, and shrubland

Diets: Insects, leaves, and shoots
Nesokia
(short-tailed bandicoot rat)

Drawing of brown rat

Gray, 1842

Two species
Scattered Asia and Egypt Size range: 15 cm (6 in) long, plus 9 cm (4 in) tail (short-tailed bandicoot rat) to 26 cm (10 in) long, plus 27 cm (11 in) tail (Bunn's short-tailed bandicoot rat)

Habitats: Forest, inland wetlands, shrubland, and grassland

Diets: Grass, grain, roots, fruit, and vegetables
Nesoromys Thomas, 1922

One species
Seram Island in Indonesia Size: 11–14 cm (4–6 in) long, plus 12–14 cm (5–6 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Nilopegamys

Drawing of brown and white rat

Osgood, 1928

One species
Ethiopia
Map of range
Size: About 15 cm (6 in)

Habitats: Inland wetlands

Diet: Plant material and invertebrates
Niviventer
(white-bellied rat)

Brown and white rat

Marshall, 1976

Seventeen species
Eastern, southern, and southeastern Asia Size range: 10 cm (4 in) long, plus 15 cm (6 in) tail (chestnut white-bellied rat) to 21 cm (8 in) long, plus 19 cm (7 in) tail (Chinese white-bellied rat)

Habitats: Forest and shrubland

Diets: Plant material and invertebrates
Notomys
(hopping mouse)

Brown mouse

Lesson, 1842

Ten species
Australia Size range: 7 cm (3 in) long, plus 11 cm (4 in) tail (dusky hopping mouse) to 13 cm (5 in) long, plus 16 cm (6 in) tail (Mitchell's hopping mouse)

Habitats: Savanna, grassland, desert, forest, rocky areas, and shrubland

Diets: Berries, leaves, seeds, and other vegetation
Oenomys
(rufous-nosed rat)

Brown rat

Thomas, 1904

Two species
Central and western Africa Size range: 13–19 cm (5–7 in) long, plus 15–20 cm (6–8 in) tail (common rufous-nosed rat)

Habitats: Savanna, forest, grassland, and shrubland

Diets: Vegetation, as well as insects
Otomys
(vlei rat)

Brown rat

F. Cuvier, 1824

28 species
Sub-Saharan Africa Size range: 11 cm (4 in) long, plus 8 cm (3 in) tail (Fynbos vlei rat) to 22 cm (9 in) long, plus 13 cm (5 in) tail (Angolan vlei rat)

Habitats: Savanna, grassland, forest, inland wetlands, rocky areas, and shrubland

Diets: Grass, semiaquatic plants, and shoots, as well as grain, seeds, berries, roots, and bark
Palawanomys Musser & Newcomb, 1983

One species
Philippines
Map of range
Size: 13–16 cm (5–6 in) long, plus 14–17 cm (6–7 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Papagomys

Dead brown rat

Sody, 1941

One species
Indonesia Size: 27–43 cm (11–17 in) long, plus 26–36 cm (10–14 in) tail

Habitats: Forest

Diet: Unknown
Parahydromys Poche, 1906

One species
New Guinea
Map of range
Size: 21–23 cm (8–9 in) long, plus 23–27 cm (9–11 in) tail

Habitats: Inland wetlands and forest

Diet: Insects and other invertebrates
Paraleptomys
(short-haired water rat)
Tate & Archbold, 1941

Two species
New Guinea
Map of range
Size range: 10 cm (4 in) long, plus 12 cm (5 in) tail (short-haired water rat) to 14 cm (6 in) long, plus 15 cm (6 in) tail (northern water rat)

Habitats: Forest

Diets: Plant material and invertebrates
Paramelomys
(long-nosed mosaic-tailed rat)
Rümmler, 1936

New Guinea and nearby islands Size range: 10 cm (4 in) long, plus 9 cm (4 in) tail (mountain mosaic-tailed rat) to 20 cm (8 in) long, plus 13 cm (5 in) tail (long-nosed paramelomys)

Habitats: Savanna and forest

Diets: Fruit, berries, and other vegetation
Parotomys
(whistling rat)

Brown rat

Thomas, 1918

Two species
Southern Africa Size range: 12–17 cm (5–7 in) long, plus 8–11 cm (3–4 in) tail (Littledale's whistling rat)

Habitats: Desert and shrubland

Diets: Grass, seeds, and shoots
Paucidentomys Rowe, Achmadi, & Esselstyn, 2012

One species
Indonesia
Map of range
Size: 15–17 cm (6–7 in) long, plus 19–20 cm (7–8 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Paulamys Musser, 1986

One species
Indonesia Size: About 16 cm (6 in)

Habitats: Forest

Diet: Insects, snails, worms, fungi, and fruit
Pelomys
(groove-toothed swamp rat)
Peters, 1852

Five species
Central Africa Size range: 10 cm (4 in) long, plus 10 cm (4 in) tail (least groove-toothed swamp rat) to 17 cm (7 in) long, plus 15 cm (6 in) tail (bell groove-toothed swamp rat)

Habitats: Savanna, inland wetlands, and grassland

Diets: Grass, swamp vegetation, and grains
Phloeomys
(slender-tailed cloud rat)

Gray and black rat

Waterhouse, 1839

Two species
Philippines Size range: 39 cm (15 in) long, plus 32 cm (13 in) tail (northern Luzon giant cloud rat) to 44 cm (17 in) long, plus 31 cm (12 in) tail (southern giant slender-tailed cloud rat)

Habitats: Forest and shrubland

Diets: Vegetation
Pithecheir
(tree rat)
Lesson, 1840

Two species
Malaysia and Indonesia Size range: 12–18 cm (5–7 in) long, plus 15–22 cm (6–9 in) tail (Malayan tree rat)

Habitats: Forest

Diets: Vegetation and insects
Pithecheirops Emmons, 1993

One species
Eastern Malaysia Size: About 11 cm (4 in)

Habitats: Forest

Diet: Plant material and invertebrates
Pogonomelomys
(lowland brush mouse)

Drawing of brown and white mouse

Rümmler, 1936

Three species
New Guinea Size range: 13 cm (5 in) long, plus 15 cm (6 in) tail (Shaw Mayer's brush mouse) to 18 cm (7 in) long, plus 20 cm (8 in) tail (grey pogonomelomys)

Habitats: Forest

Diets: Plant material and invertebrates
Pogonomys
(prehensile-tailed rat)

Gray mouse

A. Milne-Edwards, 1877

Five species
Northern Australia and New Guinea and nearby islands Size range: 9 cm (4 in) long, plus 12 cm (5 in) tail (chestnut tree mouse) to 20 cm (8 in) long, plus 26 cm (10 in) tail (D'Entrecasteaux Archipelago pogonomys)

Habitats: Forest

Diets: Leaves and grass and bamboo shoots
Praomys
(soft-furred mouse)

Gray mouse

Thomas, 1915

Seventeen species
Sub-Saharan Africa Size range: 8 cm (3 in) long, plus 10 cm (4 in) tail (least soft-furred mouse) to 15 cm (6 in) long, plus 18 cm (7 in) tail (forest soft-furred mouse)

Habitats: Savanna, grassland, inland wetlands, forest, and rocky areas

Diets: Fruit, seeds, vegetation, and insects
Protochromys Menzies, 1996

One species
Papua New Guinea Size: 13–18 cm (5–7 in) long, plus 18–21 cm (7–8 in) tail

Habitats: Forest

Diet: Fruit, berries, and other vegetation
Pseudohydromys
(moss mouse)
Rümmler, 1934

Twelve species
New Guinea Size range: 7 cm (3 in) long, plus 7 cm (3 in) tail (Laurie's moss mouse) to 12 cm (5 in) long, plus 10 cm (4 in) tail (western shrew mouse)

Habitats: Forest, grassland, and shrubland

Diets: Insects
Pseudomys
(pebble-mound mouse)

Brown mouse

Gray, 1832

23 species
Australia and Papua New Guinea Size range: 5 cm (2 in) long, plus 7 cm (3 in) tail (Bolam's mouse) to 16 cm (6 in) long, plus 14 cm (6 in) tail (Hastings River mouse)

Habitats: Savanna, grassland, desert, forest, inland wetlands, rocky areas, and shrubland

Diets: Seeds, roots, other vegetation, and insects
Rattus
(rat)

Gray rat

Fischer von Waldheim, 1803

68 species
Europe, Asia, Africa, and Australia Size range: 6 cm (2 in) long, plus 7 cm (3 in) tail (dusky rat) to 29 cm (11 in) long, plus 25 cm (10 in) tail (brown rat)

Habitats: Coastal marine, savanna, grassland, desert, intertidal marine, inland wetlands, forest, rocky areas, and shrubland

Diets: A wide variety of plant, insects, and animals
Rhabdomys
(four-striped grass rat)

Brown rat

Thomas, 1916

Four species
Sub-Saharan Africa Size range: 9–14 cm (4–6 in) long, plus 9–13 cm (4–5 in) tail (four-striped grass mouse)

Habitats: Savanna, grassland, and shrubland

Diets: Roots, seeds, berries, and grains, as well as insects and eggs
Rhynchomys
(shrewlike rat)

Drawing of brown rats

Thomas, 1895

Four species
Philippines Size range: 16 cm (6 in) long, plus 12 cm (5 in) tail (Tapulao shrew-rat) to 20 cm (8 in) long, plus 17 cm (7 in) tail (Mount Data shrew-rat)

Habitats: Forest

Diets: Insects and worms
Saxatilomys

Gray rat

Musser, 2005

One species
Indonesia
Map of range
Size: 15–17 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail

Habitats: Forest and rocky areas

Diet: Plant material and invertebrates
Solomys
(naked-tailed rat)
Thomas, 1922

Four species
Solomon Islands and Papua New Guinea Size range: 18 cm (7 in) long, plus 19 cm (7 in) tail (Isabel naked-tailed rat) to 33 cm (13 in) long, plus 25 cm (10 in) tail (Bougainville naked-tailed rat)

Habitats: Forest

Diets: Coconuts and nuts
Sommeromys Musser & Durden, 2002

One species
Indonesia Size: 9–11 cm (4 in) long, plus 17–20 cm (7–8 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Soricomys
(shrew mouse)

Brown mouse on stamp

Balete, Rickart, Heaney, Alviola, Duya, Duya, Sosa, & Jansa, 2012

Four species
Philippines Size range: 9–12 cm (4–5 in) long, plus 8–10 cm (3–4 in) tail (Co's shrew mouse)

Habitats: Forest

Diets: Worms and soft-bodied invertebrates
Srilankamys Musser, 1981

One species
Sri Lanka
Map of range
Size: About 15 cm (6 in)

Habitats: Forest

Diet: Plant material and invertebrates
Stenocephalemys
(narrow-headed rat)

Brown rat

Frick, 1914

Four species
Ethiopia Size range: 10–20 cm (4–8 in) long, plus 11–18 cm (4–7 in) tail (Ethiopian narrow-headed rat)

Habitats: Forest, grassland, and shrubland

Diets: Insects, leaves, and shoots
Stochomys Thomas, 1926

One species
Central Africa Size: 10–17 cm (4–7 in) long, plus 18–23 cm (7–9 in) tail

Habitats: Forest

Diet: Fruit, as well as vegetation and insects
Sundamys
(giant Sunda rat)
Musser & Newcomb, 1983

Three species
Indonesia and Malaysia, Southeastern Asia, and Indonesia Size range: 18–30 cm (7–12 in) long, plus 21–37 cm (8–15 in) tail (Müller's giant Sunda rat)

Habitats: Forest and shrubland

Diets: Fruit, leaves, shoots, insects, crabs, snails, and lizards
Taeromys
(Sulawesi rat)

Drawing of brown and white rats

Sody, 1841

Eight species
Indonesia Size range: 18 cm (7 in) long, plus 15 cm (6 in) tail (Sulawesi forest rat) to 28 cm (11 in) long, plus 33 cm (13 in) tail (giant Sulawesi rat)

Habitats: Forest

Diets: Fruit; Fruit, leaves, and insects
Tarsomys
(long-footed rat)
Mearns, 1905

Two species
Philippines Size range: 13 cm (5 in) long, plus 11 cm (4 in) tail (long-footed rat) to 18 cm (7 in) long, plus 16 cm (6 in) tail (spiny long-footed rat)

Habitats: Forest

Diets: Invertebrates
Tateomys
(long-tailed shrew rat)
Musser, 1969

Two species
Indonesia Size range: 11 cm (4 in) long, plus 16 cm (6 in) tail (long-tailed shrew rat) to 16 cm (6 in) long, plus 17 cm (7 in) tail (Tate's shrew rat)

Habitats: Forest

Diets: Worms
Thallomys
(acacia rat)

Gray rats

Thomas, 1920

Four species
Sub-Saharan Africa Size range: 11–17 cm (4–7 in) long, plus 12–20 cm (5–8 in) tail (black-tailed tree rat)

Habitats: Savanna, forest, and shrubland

Diets: Buds, leaves, and seeds, as well as berries, roots, and insects
Thamnomys
(thicket rat)
Thomas, 1907

Four species
Central Africa Size range: 11 cm (4 in) long, plus 17 cm (7 in) tail (Schouteden's thicket rat) to 18 cm (7 in) long, plus 23 cm (9 in) tail (Kemp's thicket rat)

Habitats: Forest

Diets: Leaves and seeds
Tokudaia
(Ryūkyū spiny rat)

Stuffed brown rat

Kuroda, 1943

Three species
Southern islands in Japan Size range: 10 cm (4 in) long, plus 8 cm (3 in) tail (Ryukyu spiny rat) to 18 cm (7 in) long, plus 14 cm (6 in) tail (Muennink's spiny rat)

Habitats: Forest

Diets: Plant material and invertebrates
Tonkinomys Musser, Lunde, & Son, 2006

One species
Vietnam Size: 18–22 cm (7–9 in) long, plus 15–19 cm (6–7 in) tail

Habitats: Forest, inland wetlands, and rocky areas

Diet: Plant material and invertebrates
Tryphomys Miller, 1910

One species
Philippines
Map of range
Size: 17–20 cm (7–8 in) long, plus 15–19 cm (6–7 in) tail

Habitats: Forest and grassland

Diet: Plant material and invertebrates
Uromys
(giant rat)

Brown and white rat

Peters, 1867

Eleven species
Southeastern Asia and northern Australia Size range: 17 cm (7 in) long, plus 18 cm (7 in) tail (masked white-tailed rat) to 38 cm (15 in) long, plus 36 cm (14 in) tail (giant white-tailed rat)

Habitats: Inland wetlands, forest, and caves

Diets: Coconuts, nuts, fruit, and flowers
Vandeleuria
(long-tailed climbing mouse)

Brown and white mouse

Gray, 1842

Three species
Southern and southeastern Asia Size range: 7 cm (3 in) long, plus 11 cm (4 in) tail (Asiatic long-tailed climbing mouse) to 13 cm (5 in) long, plus 13 cm (5 in) tail (Nolthenius's long-tailed climbing mouse)

Habitats: Forest, grassland, and shrubland

Diets: Fruit, buds, and shoorts
Vernaya Anthony, 1941

One species
China
Map of range
Size: 5–8 cm (2–3 in) long, plus 10–14 cm (4–6 in) tail

Habitats: Forest

Diet: Plant material and invertebrates
Waiomys

Brown rat

Rowe, Achmadi, & Esselstyn, 2014

One species
Indonesia Size: About 13 cm (5 in)

Habitats: Forest and inland wetlands

Diet: Plant material and invertebrates
Xenuromys Tate & Archbold, 1941

One species
New Guinea Size: 27–34 cm (11–13 in) long, plus 22–28 cm (9–11 in) tail

Habitats: Forest and rocky areas

Diet: Fruit, seeds, and insects
Xeromys

Gray mouse

Thomas, 1889

One species
Papua New Guinea and northern and eastern Australia Size: 7–13 cm (3–5 in) long, plus 6–10 cm (2–4 in) tail

Habitats: Intertidal marine, inland wetlands, coastal marine, and forest

Diet: Insects, fish, lizards, and crabs
Zelotomys
(broad-headed mouse)

Brown mouse

Osgood, 1910

Two species
Central and southern Africa Size range: 9–16 cm (4–6 in) long, plus 7–14 cm (3–6 in) tail (Woosnam's broad-headed mouse)

Habitats: Inland wetlands, savanna, forest, and grassland

Diets: Grain, as well as insects
Zyzomys
(rock rat)

Gray rat

Thomas, 1909

Five species
Australia Size range: 8 cm (3 in) long, plus 9 cm (4 in) tail (common rock rat) to 17 cm (7 in) long, plus 15 cm (6 in) tail (Arnhem Land rock rat)

Habitats: Savanna, grassland, forest, rocky areas, and shrubland

Diets: Fruit, seeds, other vegetation, and insects
Family Nesomyidae

Members of the Nesomyidae family are called nesomyids and include Malagasy rodents, climbing mice, African rock mice, pouched rats, and tufted-tailed rats. They are generally omnivores, and eat seeds, fruit, nuts, roots, stems, and insects. Nesomyidae comprises 67 extant species divided into 21 genera, grouped into six subfamilies: Cricetomyinae, containing 8 species of hamster-rats and pouched rats; Delanymyinae, containing a single species; Dendromurinae, containing 26 species of climbing mice and fat mice; Mystromyinae, containing a single species; Nesomyinae, containing 27 species of short-tailed rats, tufted-tailed rats, and big-footed mice; and Petromyscinae, containing 4 species of rock mice.

Subfamily Cricetomyinae – Roberts, 1951 – three genera
Name Authority and species Range Size and ecology
Beamys
(hamster-rat)

Gray rat

Thomas, 1909

Two species
Southeastern Africa Size range: 12 cm (5 in) long, plus 11 cm (4 in) tail (lesser hamster-rat) to 18 cm (7 in) long, plus 15 cm (6 in) tail (greater hamster-rat)

Habitats: Forest

Diets: Seeds and fruit
Cricetomys
(pouched rat)

Gray rat

Waterhouse, 1840

Four species
Central, western, and southern Africa Size range: 28 cm (11 in) long, plus 25 cm (10 in) tail (Gambian pouched rat) to 41 cm (16 in) long, plus 45 cm (18 in) tail (southern giant pouched rat)

Habitats: Forest and savanna

Diets: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails
Saccostomus

Gray mouse

Peters, 1846

Two species
Eastern and southern Africa Size range: 9–18 cm (4–7 in) long, plus 2–6 cm (1–2 in) tail (South African pouched mouse)

Habitats: Savanna, grassland, desert, and shrubland

Diets: Seeds, berries, grain, and nuts, as well as insects
Subfamily Delanymyinae – Musser & Carleton, 2005 – one genus
Name Authority and species Range Size and ecology
Delanymys Hayman, 1962

One species
Central Africa
Map of range
Size: 5–7 cm (2–3 in) long, plus 8–12 cm (3–5 in) tail

Habitats: Shrubland and inland wetlands

Diet: Seeds
Subfamily Dendromurinae – Alston, 1876 – six genera
Name Authority and species Range Size and ecology
Dendromus
(climbing mouse)

Brown mouse

Smith, 1829

Fourteen species
Sub-Saharan Africa Size range: 4 cm (2 in) long, plus 6 cm (2 in) tail (chestnut climbing mouse) to 10 cm (4 in) long, plus 9 cm (4 in) tail (Lovat's climbing mouse)

Habitats: Shrubland, forest, savanna, desert, grassland, and inland wetlands

Diets: Seeds, berries, insects, eggs, small lizards, and small birds
Dendroprionomys Petter, 1966

One species
Republic of the Congo Size: About 8 cm (3 in)

Habitats: Unknown

Diet: Insects
Malacothrix Candolle, 1838

One species
Southern Africa Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail

Habitats: Desert, shrubland, and savanna

Diet: Green vegetation
Megadendromus Dieterlen & Rupp, 1978

One species
Ethiopia Size: 11–13 cm (4–5 in) long, plus 9–11 cm (4 in) tail

Habitats: Shrubland

Diet: Likely vegetation and insects
Prionomys

Stamp with gray mouse

Dollman, 1910

One species
West-central Africa Size: 7–9 cm (3–4 in) long, plus 10–12 cm (4–5 in) tail

Habitats: Forest

Diet: Insects
Steatomys
(fat mouse)

Brown mouse

Peters, 1846

Eight species
Sub-Saharan Africa Size range: 5 cm (2 in) long, plus 3 cm (1 in) tail (tiny fat mouse) to 13 cm (5 in) long, plus 8 cm (3 in) tail (Bocage's African fat mouse)

Habitats: Shrubland, forest, savanna, and grassland

Diets: Seeds, grass bulbs, and insects
Subfamily Mystromyinae – Vorontsov, 1966 – one genus
Name Authority and species Range Size and ecology
Mystromys Wagner, 1841

One species
Southern Africa Size: 11–20 cm (4–8 in) long, plus 4–9 cm (2–4 in) tail

Habitats: Shrubland and grassland

Diet: Seeds, vegetable matter, and insects
Subfamily Nesomyinae – Forsyth Major, 1897 – nine genera
Name Authority and species Range Size and ecology
Brachytarsomys
(short-tailed rat)

Drawing of brown rodent

Günther, 1875

Two species
Northern and eastern Madagascar Size range: 22 cm (9 in) long, plus 22 cm (9 in) tail (white-tailed antsangy) to 24 cm (9 in) long, plus 27 cm (11 in) tail (hairy-tailed antsangy)

Habitats: Forest

Diets: Fruit
Brachyuromys Forsyth Major, 1896

Two species
Eastern Madagascar Size range: 14–19 cm (6–7 in) long, plus 7–10 cm (3–4 in) tail (Betsileo short-tailed rat)

Habitats: Forest, grassland, inland wetlands, and shrubland

Diets: Likely vegetation and insects
Eliurus
(tufted-tailed rat)

Brown rat

A. Milne-Edwards, 1885

Madagascar Size range: 10 cm (4 in) long, plus 11 cm (4 in) tail (lesser tufted-tailed rat) to 17 cm (7 in) long, plus 18 cm (7 in) tail (Grandidier's tufted-tailed rat)

Habitats: Forest and shrubland

Diets: Likely vegetation and insects
Gymnuromys Forsyth Major, 1896

One species
Eastern Madagascar
Map of range
Size: 15–18 cm (6–7 in) long, plus 14–20 cm (6–8 in) tail

Habitats: Forest

Diet: Likely vegetation and insects
Hypogeomys

Gray rat

Grandidier, 1869

One species
Western Madagascar
Map of range
Size: 30–34 cm (12–13 in) long, plus 21–24 cm (8–9 in) tail

Habitats: Forest

Diet: Fruit
Macrotarsomys
(big-footed mouse)

Gray mouse

A. Milne-Edwards & Grandidier, 1898

Three species
Madagascar Size range: 8 cm (3 in) long, plus 12 cm (5 in) tail (bastard big-footed mouse) to 16 cm (6 in) long, plus 24 cm (9 in) tail (Petter's big-footed mouse)

Habitats: Forest and shrubland

Diets: Berries, fruit, seeds, roots, and stems
Monticolomys Goodman & Carleton, 1996

One species
Madagascar
Map of range
Size: 9–11 cm (4 in) long, plus 13–15 cm (5–6 in) tail

Habitats: Forest

Diet: Likely vegetation and insects
Nesomys
(red forest rat)

Brown mouse

Peters, 1870

Three species
Western and eastern Madagascar Size range: 17–20 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail (island mouse)

Habitats: Forest

Diets: Fruit and nuts
Voalavo
(voalavo)
Goodman & Carleton, 1998

Two species
Central and northern Madagascar
Map of range
Size range: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail (eastern voalavo)

Habitats: Forest

Diets: Likely vegetation and insects
Subfamily Petromyscinae – Roberts, 1951 – one genus
Name Authority and species Range Size and ecology
Petromyscus
(rock mouse)
Thomas, 1926

Four species
Southern Africa Size range: 6–10 cm (2–4 in) long, plus 6–9 cm (2–4 in) tail (Brukkaros pygmy rock mouse)

Habitats: Rocky areas and shrubland

Diets: Omnivorous
Family Platacanthomyidae

Members of the Platacanthomyidae family are called platacanthomyids or Oriental dormice. They generally eat fruit, seeds, grain, and roots. Platacanthomyidae comprises two extant species in two genera.

Not assigned to a named subfamily – two genera
Name Authority and species Range Size and ecology
Platacanthomys

Brown and white dormouse

Blyth, 1859

One species
Southern India
Map of range
Size: 11–14 cm (4–6 in) long, plus 7–11 cm (3–4 in) tail

Habitats: Forest

Diet: Fruit, seeds, grain, and roots
Typhlomys

Black dormouse

Milne-Edwards, 1877

One species
Eastern China and northern Vietnam
Map of range
Size: 7–8 cm (3 in) long, plus 9–11 cm (4 in) tail

Habitats: Forest

Diet: Vegetation
Family Spalacidae

Members of the Spalacidae family are called spalacids and include blind mole-rats, bamboo rats, mole-rats, and zokors. They are primarily herbivores, and eat roots, bulbs, tubers, grass, and seeds, with some species also eating insects. Spalacidae comprises 23 extant species in 7 genera, divided into 3 subfamilies: Myospalacinae contains 6 species of zokors, Rhizomyinae contains 6 species of bamboo rats and mole-rats, Spalacinae contains 11 species of blind mole-rats.

Subfamily Myospalacinae – Lilljeborg, 1866 – two genera
Name Authority and species Range Size and ecology
Eospalax
(Chinese zokor)

Gray zokor in a cage

Allen, 1938

Three species
Central China Size range: 14 cm (6 in) long, plus 2 cm (0.8 in) tail (Rothschild's zokor) to 25 cm (10 in) long, plus 4 cm (2 in) tail (Smith's zokor)

Habitats: Shrubland, grassland, and forest

Diets: Roots and grains
Myospalax
(zokor)

Brown zokor

Laxmann, 1769

Three species
Central Asia Size range: 16 cm (6 in) long, plus 3 cm (1 in) tail (false zokor) to 27 cm (11 in) long, plus 5 cm (2 in) tail (Siberian zokor)

Habitats: Shrubland, grassland, forest, and desert

Diets: Roots and grains
Subfamily Rhizomyinae – Winge, 1887 – three genera
Name Authority and species Range Size and ecology
Cannomys

Brown bamboo rat

Thomas, 1915

One species
Southeastern Asia
Map of range
Size: 14–26 cm (6–10 in) long, plus 4–10 cm (2–4 in) tail

Habitats: Forest

Diet: Shrubs, shoots, and roots
Rhizomys
(bamboo rat)

Taxidermied brown bamboo rat

Gray, 1831

Three species
Southern and southeastern Asia Size range: 22 cm (9 in) long, plus 5 cm (2 in) tail (Chinese bamboo rat) to 48 cm (19 in) long, plus 20 cm (8 in) tail (large bamboo rat)

Habitats: Grassland and forest

Diets: Bamboo roots, as well as grass, seeds, and fruit
Tachyoryctes
(African mole-rat)

Brown mole-rat

Rüppell, 1835

Two species
Eastern Africa Size range: 15 cm (6 in) long, plus 4 cm (2 in) tail (northeast African mole-rat) to 31 cm (12 in) long, plus 7 cm (3 in) tail (giant root-rat)

Habitats: Shrubland, grassland, savanna, and forest

Diets: Roots, rhizomes, tubers, bulbs, and corms, as well as grass and legumes
Subfamily Spalacinae – Gray, 1821 – two genera
Name Authority and species Range Size and ecology
Nannospalax
(small-bodied mole-rat)

Gray mole-rat

Palmer, 1903

Three species
Southeastern Europe, Western Asia, and northeastern Africa Size range: 13 cm (5 in) long, plus 0 cm (0 in) tail (Middle East blind mole-rat) to 25 cm (10 in) long (Anatolian blind mole-rat)

Habitats: Shrubland, grassland, and desert

Diets: Roots, tubers, acorns, plant stems, and other plant parts
Spalax
(blind mole-rat)

Gray mole-rat

Güldenstädt, 1770

Eight species
Eastern Europe, Kazakhstan, and southwestern Russia Size range: 19 cm (7 in) long, plus 0 cm (0 in) tail (greater blind mole-rat) to 35 cm (14 in) long (giant blind mole-rat)

Habitats: Shrubland, grassland, and forest

Diets: Roots, bulbs, tubers, grass, and seeds, as well as insects

Suborder Sciuromorpha

Family Aplodontiidae

Members of the Aplodontiidae family are called aplodontiids. It includes a single herbivorous species.

Not assigned to a named subfamily – one genus
Name Authority and species Range Size and ecology
Aplodontia

Gray beaver

Richardson, 1829

One species
Western United States and southwestern Canada
Map of range
Size: 23–43 cm (9–17 in) long, plus 2–6 cm (1–2 in) tail

Habitats: Forest, shrubland, and inland wetlands

Diet: A wide variety of vegetation

Family Gliridae

Members of the Gliridae family are called glirids or dormice. They are omnivores and feed on fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents. Gliridae comprises 29 extant species in 9 genera, divided into 3 subfamilies: Glirinae, containing two species; Graphiurinae, containing fifteen species; and Leithiinae, containing twelve species.

Subfamily Glirinae – Muirhead, 1819 – two genera
Name Authority and species Range Size and ecology
Glirulus

Brown and white dormice

Thomas, 1906

One species
Japan
Map of range
Size: 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail

Habitats: Forest

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Glis

Brown and white dormouse

Brisson, 1762

One species
Europe and western Asia
Map of range
Size: 13–19 cm (5–7 in) long, plus 10–18 cm (4–7 in) tail

Habitats: Forest and shrubland

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Subfamily Graphiurinae – Winge, 1887 – one genus
Name Authority and species Range Size and ecology
Graphiurus
(African dormouse)

Brown and white dormouse

Smuts, 1832

Fifteen species
Sub-Saharan Africa Size range: 6 cm (2 in) long, plus 6 cm (2 in) tail (Johnston's African dormouse) to 16 cm (6 in) long, plus 13 cm (5 in) tail (Monard's dormouse)

Habitats: Grassland, inland wetlands, rocky areas, savanna, shrubland, and forest

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Subfamily Leithiinae – Lydekker, 1896 – six genera
Name Authority and species Range Size and ecology
Chaetocauda Wang, 1985

One species
Central China Size: 9–10 cm (4 in) long, plus 9–11 cm (4 in) tail

Habitats: Forest

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Dryomys
(forest dormouse)

Brown and white dormouse

Thomas, 1905

Three species
Eastern Europe and western and central Asia Size range: 7 cm (3 in) long, plus 6 cm (2 in) tail (forest dormouse) to 11 cm (4 in) long, plus 9 cm (4 in) tail (Balochistan forest dormouse)

Habitats: Shrubland, forest, and rocky areas

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Eliomys
(garden dormouse)

Brown and white dormouse

Wagner, 1840

Three species
Europe, western Asia, and northern Africa
Map of range
Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (garden dormouse) to 15 cm (6 in) long, plus 14 cm (6 in) tail (Asian garden dormouse)

Habitats: Desert, rocky areas, coastal marine, shrubland, and forest

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Muscardinus

Brown dormouse

Kaup, 1829

One species
Europe and western Asia
Map of range
Size: 6–10 cm (2–4 in) long, plus 5–9 cm (2–4 in) tail

Habitats: Forest

Diet: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Myomimus
(mouse-tailed dormouse)
Ogniov, 1924

Three species
Western Asia and southeastern Europe Size range: 6 cm (2 in) long, plus 6 cm (2 in) tail (Setzer's mouse-tailed dormouse) to 14 cm (6 in) long, plus 10 cm (4 in) tail (Roach's mouse-tailed dormouse)

Habitats: Shrubland, forest, and savanna

Diets: Fruit and nuts, as well as invertebrates, birds and their eggs, and small rodents
Selevinia

Brown and white dormouse on stamp

Belosludov & Bazhanov, 1939

One species
Kazakhstan Size: 7–10 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail

Habitats: Desert

Diet: Insects and spiders

Family Sciuridae

Members of the Sciuridae family are called sciurids and include squirrels, chipmunks, and marmots. They primarily feed on fruit, seeds, and nuts, though many species also eat insects, fungi, bark, eggs, and small vertebrates. Sciuridae comprises 284 extant species in 60 genera, divided into 5 subfamilies: Callosciurinae, containing 67 species of Asian squirrels; Ratufinae, containing 4 species of giant squirrels; Sciurillinae, containing a single species of pygmy squirrel; Sciurinae, containing 84 species of flying squirrel and tree squirrels; and Xerinae, containing 128 species of marmots, chipmunks, prairie dogs, and ground squirrels.

Subfamily Callosciurinae – Pocock, 1923 – fourteen genera
Name Authority and species Range Size and ecology
Callosciurus
(beautiful squirrel)

Gray squirrel

Gray, 1867

Fifteen species
Southeastern and eastern Asia Size range: 14 cm (6 in) long, plus 13 cm (5 in) tail (Borneo black-banded squirrel) to 29 cm (11 in) long, plus 21 cm (8 in) tail (Inornate squirrel)

Habitats: Shrubland, inland wetlands, and forest

Diets: Fruit, nuts, seeds, flowers, vegetation, insects, and eggs
Dremomys
(red-cheeked squirrel)

Brown squirrel

Heude, 1898

Six species
Southern, southeastern, and eastern Asia Size range: 15 cm (6 in) long, plus 9 cm (4 in) tail (Bornean mountain ground squirrel) to 22 cm (9 in) long, plus 17 cm (7 in) tail (red-throated squirrel)

Habitats: Shrubland, rocky areas, and forest

Diets: Fruit, nuts, vegetation, and insects
Exilisciurus
(pygmy squirrel)

Brown squirrel

Moore, 1958

Three species
Southeastern Asia Size range: 7 cm (3 in) long, plus 4 cm (2 in) tail (least pygmy squirrel) to 9 cm (4 in) long, plus 7 cm (3 in) tail (Philippine pygmy squirrel)

Habitats: Forest

Diets: Vegetation and insects
Funambulus
(palm squirrel)

Gray and white squirrel

Lesson, 1835

Six species
Southern Asia Size range: 10 cm (4 in) long, plus 7 cm (3 in) tail (dusky striped squirrel) to 16 cm (6 in) long, plus 15 cm (6 in) tail (jungle palm squirrel)

Habitats: Shrubland, grassland, inland wetlands, and forest

Diets: Seeds, nuts, bark, buds, leaves, flowers, and insects
Glyphotes

Brown squirrel

Thomas, 1898

One species
Borneo Size: 9–15 cm (4–6 in) long, plus 9–11 cm (4 in) tail

Habitats: Forest

Diet: Fruit, nuts, seeds, flowers, vegetation, insects, and eggs
Hyosciurus Archbold & Tate, 1935

Two species
Indonesia Size range: 19 cm (7 in) long, plus 6 cm (2 in) tail (Montane long-nosed squirrel) to 25 cm (10 in) long, plus 13 cm (5 in) tail (Lowland long-nosed squirrel)

Habitats: Shrubland and forest

Diets: Fruit, nuts, and insects
Lariscus

Drawing of brown squirrel

Thomas & Wroughton, 1909

Four species
Southeastern Asia Size range: 18 cm (7 in) long, plus 8 cm (3 in) tail (Niobe ground squirrel) to 20 cm (8 in) long, plus 9 cm (4 in) tail (Mentawai three-striped squirrel)

Habitats: Shrubland and forest

Diets: Fruit and nuts
Menetes

Brown squirrel

Thomas, 1908

One species
Southeastern Asia
Map of range
Size: About 18 cm (7 in)

Habitats: Grassland, shrubland, and forest

Diet: Fruit and nuts
Nannosciurus

Brown squirrels

Trouessart, 1880

One species
Southeastern Asia Size: About 8 cm (3 in)

Habitats: Forest

Diet: Fruit and nuts
Prosciurillus

Gray squirrel

Ellerman, 1947

Seven species
Indonesia and Philippines Size range: 10 cm (4 in) long, plus 5 cm (2 in) tail (Celebes dwarf squirrel) to 20 cm (8 in) long, plus 18 cm (7 in) tail (Alston's squirrel)

Habitats: Forest

Diets: Fruit, nuts, bark, and resin
Rhinosciurus

Brown squirrel

Blyth, 1856

One species
Southeastern Asia
Map of range
Size: 19–21 cm (7–8 in) long, plus 11–14 cm (4–6 in) tail

Habitats: Forest

Diet: Insects and earthworms, as well as fruit
Rubrisciurus Ellerman, 1954

One species
Island of Sulawesi in Indonesia Size: 15–30 cm (6–12 in) long, plus 18–25 cm (7–10 in) tail

Habitats: Forest

Diet: Fruit and seeds
Sundasciurus
(Sunda squirrel)

Gray squirrel

Moore, 1958

Fifteen species
Southeastern Asia Size range: 11 cm (4 in) long, plus 7 cm (3 in) tail (fraternal squirrel) to 25 cm (10 in) long, plus 25 cm (10 in) tail (horse-tailed squirrel)

Habitats: Forest

Diets: Fruit and vegetation, as well as insects
Tamiops
(Asiatic striped squirrel)

Brown squirrel

Allen, 1906

Four species
Southeastern and eastern Asia Size range: 11 cm (4 in) long, plus 10 cm (4 in) tail (Cambodian striped squirrel) to 13 cm (5 in) long, plus 11 cm (4 in) tail (maritime striped squirrel)

Habitats: Shrubland, grassland, inland wetlands, and forest

Diets: Fruit, nuts, seeds, and insects
Subfamily Ratufinae – Moore, 1959 – one genus
Name Authority and species Range Size and ecology
Ratufa
(giant squirrel)

Gray squirrel

Gray, 1867

Four species
Southern and southeastern Asia Size range: 32 cm (13 in) long, plus 35 cm (14 in) tail (grizzled giant squirrel) to 45 cm (18 in) long, plus 49 cm (19 in) tail (Indian giant squirrel)

Habitats: Forest

Diets: Fruit, nuts, bark, insects, and eggs
Subfamily Sciurillinae – Moore, 1959 – one genus
Name Authority and species Range Size and ecology
Sciurillus

Drawing of brown squirrel

Thomas, 1914

One species
Northern South America
Map of range
Size: 8–12 cm (3–5 in) long, plus 8–12 cm (3–5 in) tail

Habitats: Forest

Diet: Fruit, nuts, bark, and resin
Subfamily Sciurinae – Hemprich, 1820 – 20 genera
Name Authority and species Range Size and ecology
Aeretes

Drawing of brown squirrel

Allen, 1940

One species
Central China Size: 27–35 cm (11–14 in) long, plus 27–36 cm (11–14 in) tail

Habitats: Forest

Diet: Nuts, fruit, twigs, shoots, leaves, bark and insects
Aeromys
(large black flying squirrel)

Drawing of black squirrel

Robinson & Kloss, 1915

Two species
Southeastern Asia and Indonesia and Malaysia Size range: 30 cm (12 in) long, plus 37 cm (15 in) tail (Thomas's flying squirrel) to 38 cm (15 in) long, plus 44 cm (17 in) tail (black flying squirrel)

Habitats: Forest

Diets: Fruit, nuts, leaves, and insects
Belomys

Drawing of brown squirrels

Thomas, 1908

One species
Southeastern and eastern Asia
Map of range
Size: 18–21 cm (7–8 in) long, plus 13–17 cm (5–7 in) tail

Habitats: Forest

Diet: Nuts, fruit, twigs, shoots, leaves, bark and insects
Biswamoyopterus
(Indochinese giant flying squirrel)
Saha, 1981

Two species
Laos and eastern India Size range: 40 cm (16 in) long, plus 60 cm (24 in) tail (Namdapha flying squirrel) to 46 cm (18 in) long, plus 62 cm (24 in) tail (Laotian giant flying squirrel)

Habitats: Forest

Diets: Unknown
Eoglaucomys Howell, 1915

One species
Central Asia Size: 23–30 cm (9–12 in) long, plus 25–33 cm (10–13 in) tail

Habitats: Forest

Diet: Seeds, as well as shoots, buds, leaves, and nuts
Eupetaurus

Drawing of brown squirrel

Thomas, 1888

One species
Western China Size: 42–61 cm (17–24 in) long, plus 38–54 cm (15–21 in) tail

Habitats: Forest, rocky areas, and caves

Diet: Moss and lichen
Glaucomys
(New World flying squirrel)

Brown squirrel

Thomas, 1908

Two species
Eastern North America and Canada and northern United States Size range: 11–19 cm (4–7 in) long, plus 12–16 cm (5–6 in) tail (Northern flying squirrel)

Habitats: Forest

Diets: Nuts, bark, lichen, fungi, fruit, and berries, as well as insects
Hylopetes
(arrow-tailed flying squirrel)

Gray squirrel

Thomas, 1908

Nine species
Southeastern and eastern Asia Size range: 11 cm (4 in) long, plus 9 cm (4 in) tail (arrow flying squirrel) to 33 cm (13 in) long, plus 31 cm (12 in) tail (Palawan flying squirrel)

Habitats: Shrubland and forest

Diets: Nuts, shoots, leaves, fruit, insects, and small snakes
Iomys
(Horsfield's flying squirrel)

Drawing of brown squirrel

Thomas, 1908

Two species
Southeastern Asia and Indonesia Size range: 16–23 cm (6–9 in) long, plus 15–23 cm (6–9 in) tail (Javanese flying squirrel)

Habitats: Shrubland and forest

Diets: Nuts, fruit, twigs, shoots, leaves, bark and insects
Microsciurus
(dwarf squirrel)

Gray squirrel

Allen, 1895

Four species
Central America and northern South America Size range: 10 cm (4 in) long, plus 8 cm (3 in) tail (Central American dwarf squirrel) to 16 cm (6 in) long, plus 15 cm (6 in) tail (Amazon dwarf squirrel)

Habitats: Forest

Diets: Fruit and nuts
Petaurillus
(pygmy flying squirrel)
Thomas, 1908

Three species
Borneo and Malaysia Size range: 6 cm (2 in) long, plus 6 cm (2 in) tail (lesser pygmy flying squirrel) to 10 cm (4 in) long, plus 10 cm (4 in) tail (Selangor pygmy flying squirrel)

Habitats: Forest

Diets: Nuts, fruit, twigs, shoots, leaves, bark and insects
Petaurista
(giant flying squirrel)

Brown and white squirrel

Link, 1795

Asia Size range: 27 cm (11 in) long, plus 28 cm (11 in) tail (Japanese giant flying squirrel) to 61 cm (24 in) long, plus 69 cm (27 in) tail (Indian giant flying squirrel)

Habitats: Rocky areas and forest

Diets: Fruit, nuts, twigs, shoots, and leaves, as well as insects
Petinomys
(dwarf flying squirrel)

Drawing of squirrel

Thomas, 1908

Eight species
Southeastern Asia and southern India Size range: 9 cm (4 in) long, plus 8 cm (3 in) tail (Temminck's flying squirrel) to 37 cm (15 in) long, plus 46 cm (18 in) tail (Mindanao flying squirrel)

Habitats: Forest

Diets: Nuts, fruit, twigs, shoots, and leaves, as well as bark and insects
Pteromys
(Old World flying squirrel)

Brown squirrel

Cuvier, 1800

Two species
Japan, northern Asia, and northern Europe Size range: 12–23 cm (5–9 in) long, plus 9–15 cm (4–6 in) tail (Siberian flying squirrel)

Habitats: Forest

Diets: Nuts, pine seeds, buds, bark, fruit, and insects
Pteromyscus

Drawing of gray squirrel

Thomas, 1908

One species
Southeastern Asia Size: 20–29 cm (8–11 in) long, plus 17–23 cm (7–9 in) tail

Habitats: Forest

Diet: Nuts, fruit, twigs, shoots, leaves, bark and insects
Rheithrosciurus

Brown, black, and white squirrel

Gray, 1867

One species
Borneo
Map of range
Size: 33–35 cm (13–14 in) long, plus 30–34 cm (12–13 in) tail

Habitats: Forest

Diet: Fruit and nuts
Sciurus
(tree squirrel)

Brown squirrel

Linnaeus, 1758

29 species
North America, South America, Europe, and Asia Size range: 14 cm (6 in) long, plus 11 cm (4 in) tail (Andean squirrel) to 56 cm (22 in) long, plus 27 cm (11 in) tail (Peters's squirrel)

Habitats: Shrubland, grassland, and forest

Diets: Nuts, seeds, fruit, buds, and tree shoots
Syntheosciurus

Brown squirrel

Bangs, 1902

One species
Costa Rica and Panama Size: 15–19 cm (6–7 in) long, plus 12–16 cm (5–6 in) tail

Habitats: Forest

Diet: Flowers, buds, leaves, and sap
Tamiasciurus
(American pine squirrel)

Brown squirrel

Trouessart, 1880

Three species
North America Size range: 18–19 cm (7 in) long, plus 11–15 cm (4–6 in) tail (multiple)

Habitats: Forest

Diets: Nuts, buds, fruit, bark, fungi, sap, eggs, and small vertebrates
Trogopterus

Drawing of brown squirrel

Heude, 1898

One species
Southern China Size: 20–33 cm (8–13 in) long, plus 26–30 cm (10–12 in) tail

Habitats: Forest and caves

Diet: Leaves, twigs, fruit, and nuts
Subfamily Xerinae – Osborn, 1910 – 24 genera
Name Authority and species Range Size and ecology
Ammospermophilus
(antelope squirrel)

Brown squirrel

Merriam, 1892

Four species
Western United States and western Mexico Size range: 18 cm (7 in) long, plus 4 cm (2 in) tail (white-tailed antelope squirrel) to 28 cm (11 in) long, plus 10 cm (4 in) tail (Harris's antelope squirrel)

Habitats: Shrubland, grassland, desert, and forest

Diets: Seeds, fruit, stems, and roots, as well as insects and carrion
Atlantoxerus

Brown and white squirrel

Forsyth Major, 1893

One species
Northwestern Africa
Map of range
Size: About 17 cm (7 in)

Habitats: Rocky areas, grassland, and shrubland

Diet: Fruit and seeds
Callospermophilus
(golden-mantled ground squirrel)

Brown squirrel

Merriam, 1897

Three species
Western North America Size range: 16 cm (6 in) long, plus 6 cm (2 in) tail (Sierra Madre ground squirrel) to 31 cm (12 in) long, plus 12 cm (5 in) tail (Cascade golden-mantled ground squirrel)

Habitats: Shrubland, rocky areas, grassland, and forest

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Cynomys
(prairie dog)

Brown prairie dog

Rafinesque, 1817

Five species
Southern Canada, western United States, and northern Mexico
Map of range
Size range: 29 cm (11 in) long, plus 4 cm (2 in) tail (Utah prairie dog) to 44 cm (17 in) long, plus 11 cm (4 in) tail (Mexican prairie dog)

Habitats: Shrubland, savanna, grassland, and desert

Diets: Herbs and grasses
Epixerus

Brown squirrel

Thomas, 1909

One species
Western Africa
Map of range
Size: 28–29 cm (11–11 in) long, plus about 28 cm (11 in) tail

Habitats: Forest

Diet: Fruit as well as insects
Eutamias

Brown chipmunk

Trouessart, 1880

One species
Northern and eastern Asia and Europe
Map of range
Size: 14–15 cm (6 in) long, plus 10–12 cm (4–5 in) tail

Habitats: Forest and shrubland

Diet: Nuts and seeds, as well as buds, berries, grains, and fungi
Funisciurus
(rope squirrel)

Brown squirrel

Trouessart, 1880

Ten species
Central and western Africa Size range: 16 cm (6 in) long, plus 16 cm (6 in) tail (Congo rope squirrel) to 25 cm (10 in) long, plus 20 cm (8 in) tail (Lunda rope squirrel)

Habitats: Shrubland, rocky areas, savanna, and forest

Diets: Seeds, nuts, and fruit, as well as insects and eggs
Heliosciurus
(sun squirrel)

Gray squirrel

Trouessart, 1880

Six species
Sub-Saharan Africa Size range: 18 cm (7 in) long, plus 20 cm (8 in) tail (small sun squirrel) to 24 cm (9 in) long, plus 27 cm (11 in) tail (Zanj sun squirrel)

Habitats: Savanna and forest

Diets: Seeds, nuts, fruit, insects, and eggs
Ictidomys
(lined ground squirrel)

Brown squirrel

Allen, 1877

Two species
North America Size range: 17 cm (7 in) long, plus 6 cm (2 in) tail (thirteen-lined ground squirrel) to 38 cm (15 in) long, plus 17 cm (7 in) tail (Mexican ground squirrel)

Habitats: Shrubland, grassland, and coastal marine

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Marmota
(marmot)

Brown groundhog

Blumenbach, 1779

Fourteen species
Asia, Europe, United States, and Canada Size range: 36 cm (14 in) long, plus 11 cm (4 in) tail (Tarbagan marmot) to 75 cm (30 in) long, plus 24 cm (9 in) tail (Olympic marmot)

Habitats: Rocky areas, forest, grassland, shrubland, and desert

Diets: Grass and forbs, as well as fruit, grains, legumes, and insects
Myosciurus Thomas, 1909

One species
Western Africa
Map of range
Size: 7–8 cm (3 in) long, plus 5–6 cm (2 in) tail

Habitats: Forest

Diet: Fruit, bark, and insects
Neotamias
(chipmunk)

Brown chipmunk

Howell, 1929

23 species
North America
Map of range
Size range: 10 cm (4 in) long, plus 6 cm (2 in) tail (alpine chipmunk) to 16 cm (6 in) long, plus 11 cm (4 in) tail (Durango chipmunk)

Habitats: Rocky areas, forest, grassland, shrubland, and desert

Diets: Fruit and seeds, as well as flowers, fungi, insects, and eggs
Notocitellus
(tropical ground squirrel)

Gray squirrel

Howell, 1938

Two species
Mexico Size range: 16 cm (6 in) long, plus 13 cm (5 in) tail (tropical ground squirrel) to 22 cm (9 in) long, plus 21 cm (8 in) tail (ring-tailed ground squirrel)

Habitats: Shrubland, rocky areas, and forest

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Otospermophilus
(American rock squirrel)

Gray squirrel

Brandt, 1844

Two species
Mexico and western United States Size range: 28 cm (11 in) long, plus 13 cm (5 in) tail (California ground squirrel) to 54 cm (21 in) long, plus 26 cm (10 in) tail (rock squirrel)

Habitats: Shrubland, rocky areas, and grassland

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Paraxerus
(African bush squirrel)

Gray squirrel

Forsyth Major, 1893

Eleven species
Sub-Saharan Africa Size range: 13 cm (5 in) long, plus 10 cm (4 in) tail (Alexander's bush squirrel) to 23 cm (9 in) long, plus 20 cm (8 in) tail (black and red bush squirrel)

Habitats: Shrubland, savanna, and forest

Diets: Seeds and fruit, as well as roots and eggs
Poliocitellus

Gray squirrel

Howell, 1938

One species
Central United States and southern Canada Size: 23–24 cm (9 in) long, plus about 13 cm (5 in) tail

Habitats: Savanna, grassland, and inland wetlands

Diet: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Protoxerus
(African giant squirrel)

Gray squirrel

Forsyth Major, 1893

Two species
Western and central Africa Size range: 24 cm (9 in) long, plus 30 cm (12 in) tail (slender-tailed squirrel) to 30 cm (12 in) long, plus 31 cm (12 in) tail (forest giant squirrel)

Habitats: Forest

Diets: Seeds, nuts, and fruit
Sciurotamias
(Chinese rock squirrel)

Gray squirrel

Miller, 1901

Two species
China Size range: 20 cm (8 in) long, plus 14 cm (6 in) tail (Père David's rock squirrel) to 22 cm (9 in) long, plus 16 cm (6 in) tail (Forrest's rock squirrel)

Habitats: Shrubland and rocky areas

Diets: Fruit and seeds
Spermophilopsis

Brown squirrel

Blasius, 1884

One species
Central Asia Size: 23–27 cm (9–11 in) long, plus 2–9 cm (1–4 in) tail

Habitats: Desert, grassland, and shrubland

Diet: Fruit, seeds, vegetation, and insects
Spermophilus
(Holarctic ground squirrel)

Brown squirrel

F. Cuvier, 1825

Fifteen species
Asia and eastern Europe Size range: 14 cm (6 in) long, plus 3 cm (1 in) tail (Asia Minor ground squirrel) to 32 cm (13 in) long, plus 11 cm (4 in) tail (Russet ground squirrel)

Habitats: Rocky areas, grassland, shrubland, and desert

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Tamias

Brown chipmunk

Illiger, 1811

One species
Eastern United States and eastern Canada
Map of range
Size: 14–15 cm (6 in) long, plus 8–10 cm (3–4 in) tail

Habitats: Shrubland and forest

Diet: Nuts, seeds, fruit, and fungi
Urocitellus
(ground squirrel)

Brown squirrel

Obolenskij, 1927

Twelve species
Canada, United States, and central and eastern Asia Size range: 15 cm (6 in) long, plus 3 cm (1 in) tail (Merriam's ground squirrel) to 34 cm (13 in) long, plus 9 cm (4 in) tail (Richardson's ground squirrel)

Habitats: Shrubland, grassland, desert, and forest

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Xerospermophilus
(desert ground squirrel)

Brown squirrel

Merriam, 1892

Four species
Mexico and United States Size range: 18 cm (7 in) long, plus 5 cm (2 in) tail (spotted ground squirrel) to 28 cm (11 in) long, plus 12 cm (5 in) tail (round-tailed ground squirrel)

Habitats: Grassland, desert, and forest

Diets: Seeds, nuts, grains, roots, bulbs, fungi, vegetation, and insects, as well as small vertebrates and eggs
Xerus
(unstriped ground squirrel)

Brown squirrel

Hemprich & Ehrenberg, 1833

Four species
Sub-Saharan Africa Size range: 22–29 cm (9–11 in) long, plus 18–26 cm (7–10 in) tail (African striped ground squirrel)

Habitats: Shrubland, desert, savanna, and forest

Diets: Roots, seeds, fruit, grains, insects, small vertebrates, and eggs

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  138. 1 2 3 4 5 6 7 8 9 10 11 12 13 Chernasky; Motis; Burgin, pp. 206–209
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  140. 1 2 "Allactodipus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  151. 1 2 "Euchoreutes". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
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  156. 1 2 3 Chernasky; Motis; Burgin, pp. 205–206
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  164. 1 2 3 4 5 6 7 8 9 Nowak, pp. 1419–1423
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  512. 1 2 3 4 5 6 7 Chernasky; Motis; Burgin, pp. 210–213
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  524. 1 2 3 4 5 6 7 8 9 Chernasky; Motis; Burgin, pp. 402–403
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  569. ↑ "Syntheosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  570. ↑ "Tamiasciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  571. ↑ "Trogopterus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  572. ↑ "Ammospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  573. ↑ "Atlantoxerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  574. ↑ "Callospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  575. ↑ "Cynomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  576. ↑ "Epixerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  577. ↑ "Eutamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  578. ↑ "Funisciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  579. ↑ "Heliosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  580. ↑ "Ictidomys". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  581. ↑ "Marmota". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  582. ↑ "Myosciurus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  583. ↑ "Neotamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  584. ↑ "Notocitellus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  585. ↑ "Otospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  586. ↑ "Paraxerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  587. ↑ "Poliocitellus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  588. ↑ "Protoxerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  589. ↑ "Sciurotamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  590. ↑ "Spermophilopsis". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  591. ↑ "Spermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  592. ↑ "Tamias". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  593. ↑ "Urocitellus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  594. ↑ "Xerospermophilus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.
  595. ↑ "Xerus". IUCN Red List of Threatened Species. International Union for Conservation of Nature. Retrieved January 30, 2026.

Sources